II. Man a Vertebrate.
Man is a vertebrate animal.[1] He has (anatomically at least) a backbone. He belongs to that substantial class of organisms possessing an articulating internal skeleton—the family of the fishes, amphibians, reptiles, birds, and mammals. Most animals have some sort of skeleton, some sort of calcareous contrivance, whose business it is to give form and protection to the softer parts of the organism. Some animals, as the starfishes, have plates of lime scattered throughout the surface parts of the body; others, as the corals and sponges secrete plant-like frames, upon and among the branches of which the organisms reside; and still others, as the clams, crustaceans, and insects, have skeletons consisting of a shell or sheath on the outside of, and more or less surrounding, the softer substances of the body. The limbs of insects are tiny tubes on the inside of which are the miniature muscles with which they perform their marvels of locomotion. The skeleton of vertebrates, consisting of levers, beams, columns, and arches, all skilfully joined together and sunk deep within the muscular tissue, forms a conspicuous contrast to the rudimentary frames of other animals. The vertebrate skeleton consists of a hollow axis, divided into segments and extending along the dorsal region of the body, from the ventral side of which articulate, by means of awkwardly-constructed girdles, an anterior and a posterior pair of limbs. This dorsal axis ends in front in a peculiar bulbous arrangement called the head, which contains, among other valuables, the brain and buccal cavern. The thoracic segments of the backbone send off pairs of flat bones, which, arching ventrally, form the chest for the protection of the heart and other vitals. The limbs (except in fishes) consist each of a single long bone, succeeded by two long bones, followed by two transverse rows of short, irregular wrist or ankle bones, ending normally in five branching series of bones called digits. This is essentially the skeleton of all fishes, amphibians, reptiles, birds, and mammals. In short, it is the universal vertebrate type of frame. There are minor modifications to suit the various kinds of environment, adaptations to the necessities of aquatic, terrestrial, and aerial locomotion and life, some parts being specialised, others atrophied, and still others omitted, but there is never anywhere, from fishes to philosophers, any fundamental departure from the established vertebrate type of skeleton.[2] The pectoral fins of fishes correspond to the fore-limbs of frogs and reptiles, the wings of birds, and the arms of men. The pelvic fins of fishes are homologous with the hind-limbs of frogs, reptiles, and quadrupeds, and the legs of birds, apes, and men. The foot of the dog and crocodile, the hand of the orang, and the flipper of the dolphin and seal, all have the same general structure as the hand of man; and the wings of the bat and bird, the forelimbs of the lizard and elephant, and the comical shovels of the mole and ornithorhynchus, notwithstanding the great differences in their external appearance and use, contain essentially the same bones and the same arrangement of the bones as do the arms of men and women. The human body has two primary cavities in it. So have the bodies of all vertebrates: a neural cavity containing the brain and spinal cord, and a visceral cavity containing the heart, liver, lungs, and alimentary canal. Invertebrates have only one body cavity—the one corresponding to the visceral cavity of vertebrates—and the main nerve trunk, instead of extending along the back, as among vertebrates, is in invertebrates located ventrally. Vertebrates are the only animals on the earth that have a highly developed circulatory system, a system entirely shut off from the other systems, and containing a heart, arteries, veins, and capillaries. In all invertebrates the digestive and circulatory systems remain to a greater or less extent connected, the blood and food mingling more or less in the general cavity of the body. Worms and insects have pulsating tubes instead of heart and arteries. Crustaceans have hearts with one chamber, and mollusks have two or three chambered hearts, but the blood, instead of returning to the heart after its journey through the arteries, passes into the body cavity. In man and other vertebrates the circulating current is confined strictly to the bloodvessels, no particle of it ever escaping into the general body cavity. The heart of vertebrates is distinguished from that of invertebrates by being located ventrally. The heart of invertebrates is in the back. The blood of vertebrates differs from that of invertebrates in containing both red and white corpuscles. Invertebrates have white corpuscles only. Worms have yellow, red, or bright green blood. The blood of crustaceans is bluish, that of mollusks is white, and that of insects dusky or brown. The blood of all vertebrates, excepting amphioxus, is red. All backboned beings, whether they dwell in seas or cities, and whether they build nests or empires, have two eyes, two ears, nose and mouth, all located in the head, and always occupying the same relative position to each other. Invertebrates may have their brains in their abdomen, as do the mites; hear with their legs or antennae, as many insects do; see with their tunics, like the scallops; and breathe with their skin, as do the worms. The crayfish hears with its ‘feelers,’ the cricket and katydid with their fore-legs, the grasshopper with its abdomen, the clam with its ‘foot,’ and mysis and other low crustaceans have their auditory organs on their tails.
Man is, then, like the fishes, frogs, reptiles, birds, and quadrupeds, a vertebrate animal. Excepting in his infancy, when he is a quadruped going on all fours, he uses his posterior limbs only for locomotion, and his anterior for prehension and the like. His spinal axis is erect instead of horizontal, and his tail is atrophied. But he possesses all of the unmistakable qualities of the vertebrate type of structure—a two-chambered body cavity, a highly developed and dorsally located nerve trunk, vertebrate vitals, a closed circulatory system, a ventral heart, red blood, a head containing sense organs and brain, and a well-ordered internal skeleton, consisting of a vertebral column with skull and ribs and two pairs of limbs, the limbs consisting each of one long bone, two long bones, two transverse rows of irregular bones, and five branches at the end.
[1.] See ‘Classes of Animals,’ at the end of the chapter.
[2.] Snakes are limbless, and hind-limbs are lacking in whales and other degenerates; but rudimentary limbs are found in the embryonic stages of these animals. Frogs, it may be said also, have no ribs.