The little seedlings of many, especially the conifers, are quite delicate, and remain very small the first season; they need, therefore, the protecting shade of mother trees, or artificial shading, and also protection against weeds. The amount of light or shade given requires careful regulation for some of them; too much light and heat will kill them, and so will too much shade. This accounts for the failure of many seedlings that spring up in the virgin forest.
The planter, then, is required to know the nature and the needs of the various kinds of seeds and seedlings, so as to provide favorable conditions, when he will avoid sowing in the open field such as require the care which it is impracticable to give outside of the nursery.
GROWTH IN LENGTH AND RAMIFICATION.
While the stalk of wheat or corn grows for one season, exhausts itself in seed production, and then dies, the tree continues to grow from season to season, in length as well as in thickness. The growth in length of shaft and branches proceeds from buds, made up of cell tissues, which can subdivide and lengthen into shoots, as well as make leaves. These buds are formed during summer, and when winter begins contain embryo leaves, more or less developed, under the protecting cover of scales ([fig. 3]). When spring stimulates the young plant to new activity, the buds swell, shed their scales, distend their cells, increasing their number by subdivision, and thus the leaves expand, and the bud lengthens into a shoot and twig. During the season new buds are formed, and the whole process repeats itself from year to year, giving rise to the ramification and height growth of the tree. The end buds being mostly stronger and better developed, the main axis of tree or branch increases more rapidly than the rest. All these buds originate from the youngest, central part of the shoot, the pith, and hence when the tree grows in thickness, enveloping the base of the limbs, their connection with the pith can always be traced. This is the usual manner of bud formation; in addition, so-called "adventitious" buds maybe formed from the young living wood in later life, which are not connected with the pith. Such buds are those which develop into sprouts from the stump when the tree is cut; also those which give rise to what are known as "water sprouts." Many buds, although formed, are, however, not developed at once, and perhaps not at all, especially as the tree grows older; these either die or remain "dormant," often for a hundred years, to spring into life when necessary ([fig. 4]).
The fact that each ordinary limb starts as a bud from the pith is an important one to the timber grower; it explains knotty timber and gives him the hint that in order to obtain clear timber the branches first formed must be soon removed, either by the knife or by proper shading, which kills the branches and thus "clears" the shaft.
The planter has it also in his power to influence the form development of the tree by removing some of the buds, giving thereby better chance to the remaining ones. This pruning of buds is, where practicable, often better practice than the pruning of limbs.
Since the tree does not grow in length except by its buds it is evident that a limb which started to grow at the height of 6 feet has its base always 6 feet from the ground, and if allowed to grow to size, must be surrounded by the wood which accumulates on the main stem or trunk. If a limb is killed and broken off early, only a slender stub composed entirely of rapidly decaying sapwood, is left, occasioning, therefore, only a small defect in the heart of the tree; but if left to grow to considerable age, the base of the limb is encased by the wood of the stem, which, when the tree is cut into lumber, appears as a knot. The longer the limb has been allowed to grow, the farther out is the timber knotty and the thicker is the knot. If the limb remained alive, the knot is "sound," closely grown together with the fibers of the tree. If the limb died off, the remaining stub may behave in different ways. In pines it will be largely composed of heartwood, very resinous and durable; separated from the fibers of the overgrowing wood, it forms a "loose" knot, which is apt to fall out of a board, leaving a hole.