The growing tree, in all its parts, is more or less saturated with water, and as the leaves, under the influence of sun and wind and atmospheric conditions generally transpire, new supplies are taken in through the roots and conveyed to the crown. This movement takes place even in winter, in a slight degree, to supply the loss of water by evaporation from the branches. In the growing season it is so active as to become noticeable; hence the saying that the sap is "up," or "rising," and when, toward the end of the season, the movement becomes less, the sap is said to be "down." But this movement of water is always upward; hence the notion that there is a stream upward at one season and in one part of the tree, and a stream downward at another season and perhaps in another part of the tree, is erroneous. The downward movement is of food materials, and the two movements of water upward and food downward take place simultaneously, and depend, in part at least, one upon the other, the food being carried to the young parts, wherever required, by a process of diffusion from cell to cell known as "osmosis."
Fig. 1.—Physiological importance of different parts of the tree; pathways of water and food materials. (Schematic.)
These food materials are, by the life processes of the active cells, changed in chemical composition as need be, from sugar, which is soluble, into starch, which is insoluble, and back into sugar, and combined with nitrogenous substances to make the cell-forming material, protoplasm ([fig. 1]).
In the fall, when the leaves cease to elaborate food, both the upward and the downward movement, more or less simultaneously, come to rest (the surplus of food materials, as starch, and sometimes as sugar, being stored for the winter in certain cell tissues), to begin again simultaneously when in spring the temperature is high enough to reawaken activity, when the stored food of last year is dissolved and started on its voyage. The exact manner in which this movement of water upward and food materials downward takes place, and the forces at work, are not yet fully understood, nor is there absolute certainty as to the parts of the tree in which the movement takes place. It appears, however, that while all the so-called "sapwood" is capable of conducting water (the heartwood is probably not), the most active movement of both water and food materials takes place in the cambium (the growing cells immediately beneath the bark) and youngest parts of the bark.
The deductions from these processes important to the planter are: That injury to the living bark or bast means injury to growth, if not destruction to life; that during the period of vegetation transplanting can be done only with great caution; that the best time to move trees is in the fall, when the leaves have dropped and the movement of water and food materials has mostly ceased, or in spring, before the movement begins again, the winter being objectionable only because of the difficulty of working the soil and of keeping the roots protected against frost. All things considered, spring planting, before activity in the tree has begun, is the best, although it is not impossible to plant at other times.
PROGRESS OF DEVELOPMENT.
Like the wheat or corn plant, the tree seed require as conditions for sprouting sufficient moisture, warmth, and air. Tree seeds, however, differ from grain in that most of the kinds lose their power of germination easily; with few exceptions (locust, pine, spruce), they can not be kept for any length of time.
The first leaves formed often differ essentially in shape from those of the mature tree, which may cause their being confounded with other plants, weeds, etc.