On poorer sites the rate is slower, but remains longer on the increase, while on good sites the maximum rate is soon reached.
Of course, in a forest, where light conditions are not most favorable, because form development and soil conditions require shade, the total wood formation is less than in an isolated tree, favorably placed. Just so the dominant trees in a forest—i. e., those which have their crowns above all others—show, of course, the advantage they have over the inferior trees which are suffering from the shade of their neighbors.
Finally, if we would take into consideration an entire forest growth, and determine, for instance, how much wood an acre of such forest produces at different periods, we must not overlook the fact that the number of trees per acre changes as the trees grow older. Some of them are overshaded and crowded out by the others, so that a young growth of spruce might start with 100,000 little seedlings to the acre, of which in the twentieth year only 10,000 would be alive, while in the fortieth year the number would be reduced to 1,200, and in the hundredth year to 280. Hence the rate of growth of any single tree gives no idea of what the acre of forest will do.
Tims, while a single good white pine might grow the fastest in volume when about one hundred years old, then making wood at the rate of, say, 1.5 cubic feet per year, an acre of pine on good soil, containing about 1,600 trees, may make the most wood in the thirtieth year, then growing at the rate of 170 cubic feet per acre, while in the hundredth year the rate would not exceed 70 cubic feet; and an acre of pine in a poorer location, with about 1,400 trees, may make the most wood in the fortieth year, at the rate of 100 cubic feet per acre.
From the consideration of the relation of light conditions to soil conditions, to form development, and to rate of growth, we may make the following deductions of interest to the forest planter:
In order to secure the best results in wood production, in quantity and quality, at the same time preserving favorable soil conditions, the forest should be composed of various species, a mixture of light-needing and shade-enduring kinds. The light-needing ones should be of quicker growth; the shady ones, in larger numbers, should be slower growers. For the first fifteen to twenty-five years the plantation should be kept as dense as possible, to secure clear shafts and good growth in height; then it should be thinned, to increase crown development and diameter growth; the thinning, however, is not to be so severe that the crowns can not close up again in two or three years; the thinning is to be repeated again and again, always favoring the best developed trees.
REPRODUCTION.
All trees reproduce themselves naturally from seed. Man can secure their reproduction also from cuttings or layers; and some kinds can reproduce themselves by shoots from the stump when the parent tree has been cut. This latter capacity is possessed in a varying degree by different species; chestnuts, oaks, elms, maples, poplars, and willows are most excellent sprouters; most conifers do not sprout at all, and the shoots of those that do sprout soon die (Sequoia or California redwood seems to be an exception). Sprouts of broad-leaved trees develop differently from seedlings, growing very rapidly at first, but soon lessening in the rate of growth and never attaining the height and perhaps not the diameter of trees grown from the seed; they are also shorter lived. With age the stumps lose their capacity for sprouting. To secure best results, the parent tree should be cut close to the ground in early spring, avoiding severe frost, and a sharp cut should be made which will not sever the bark from the trunk.
Not all trees bear seed every year, and plentiful seed production, especially in a forest, occurs, as a rule, periodically. The periods differ with species, climate, and season.
Not all seeds can germinate, and in some species the number of seeds that can germinate is very small, and they lose their power of germination when kept a few hours, like the willows. Others, if kept till they have become dry, will "lie over" in the soil a year or more before germinating. The same thing will occur if they are covered too deep in the soil, provided they germinate at all under such conditions.