I thank you sincerely for your letter. I am very glad to hear about Lathyrus odoratus, for here in England the vars. never cross, and yet are sometimes visited by bees. (714/1. In "Cross and Self-Fertilisation," page 156, Darwin quotes the information received from Delpino and referred to in the present letter—namely, that it is the fixed opinion of the Italian gardeners that the varieties do intercross. See Letter 709.) Pisum sativum I have also many times seen visited by Bombus. I believe the cause of the many vars. not crossing is that under our climate the flowers are self-fertilised at an early period, before the corolla is fully expanded. I shall examine this point with L. odoratus. I have read H. Muller's book, and it seems to me very good. Your criticism had not occurred to me, but is, I think just—viz. that it is much more important to know what insects habitually visit any flower than the various kinds which occasionally visit it. Have you seen A. Kerner's book "Schutzmittel des Pollens," 1873, Innsbruck. (714/2. Afterwards translated by Dr. Ogle as "Flowers and their Unbidden Guests," with a prefatory letter by Charles Darwin, 1878.) It is very interesting, but he does not seem to know anything about the work of other authors.
I have Bentham's paper in my house, but have not yet had time to read a word of it. He is a man with very sound judgment, and fully admits the principle of evolution.
I have lately had occasion to look over again your discussion on anemophilous plants, and I have again felt much admiration at your work. (714/3. "Atti della Soc. Italiana di Scienze Nat." Volume XIII.)
(714/4. In the beginning of August, 1873, Darwin paid the first of several visits to Lord Farrer's house at Abinger. When sending copies of Darwin's letters for the "Life and Letters," Lord Farrer was good enough to add explanatory notes and recollections, from which we quote the following sketch.)
"Above my house are some low hills, standing up in the valley, below the chalk range on the one hand and the more distant range of Leith Hill on the other, with pretty views of the valley towards Dorking in one direction and Guildford in the other. They are composed of the less fertile Greensand strata, and are covered with fern, broom, gorse, and heath. Here it was a particular pleasure of his to wander, and his tall figure, with his broad-brimmed Panama hat and long stick like an alpenstock, sauntering solitary and slow over our favourite walks, is one of the pleasantest of the many pleasant associations I have with the place."
LETTER 715. TO T.H. FARRER (Lord Farrer).
(715/1. The following note by Lord Farrer explains the main point of the letter, which, however, refers to the "bloom" problem as well as to Coronilla:—
"I thought I had found out what puzzled us in Coronilla varia: in most of the Papilionaceae, when the tenth stamen is free, there is nectar in the staminal tube, and the opening caused by the free stamen enables the bee to reach the nectar, and in so doing the bee fertilises the plant. In Coronilla varia, and in several other species of Coronilla, there is no nectar in the staminal tube or in the tube of the corolla. But there are peculiar glands with nectar on the outside of the calyx, and peculiar openings in the tube of the corolla through which the proboscis of the bee, whilst entering the flower in the usual way and dusting itself with pollen, can reach these glands, thus fertilising the plant in getting the nectar. On writing this to Mr. Darwin, I received the following characteristic note.
The first postscript relates to the rough ground behind my house, over which he was fond of strolling. It had been ploughed up and then allowed to go back, and the interest was to watch how the numerous species of weeds of cultivation which followed the plough gradually gave way in the struggle for existence to the well-known and much less varied flora of an English common.")
Bassett, Southampton, August 14th, 1873.