You are the man to conquer a Coronilla. (715/2. In a former letter to Lord Farrer, Darwin wrote: "Here is a maxim for you, 'It is disgraceful to be beaten by a Coronilla.'") I have been looking at the half-dried flowers, and am prepared to swear that you have solved the mystery. The difference in the size of the cells on the calyx under the vexillum right down to the common peduncle is conspicuous. The flour still adhered to this side; I see little bracteae or stipules apparently with glandular ends at the base of the calyces. Do these secrete? It seems to me a beautiful case. When I saw the odd shape of the base of the vexillum, I concluded that it must have some meaning, but little dreamt what that was. Now there remains only the one serious point—viz.the separation of the one stamen. I daresay that you are right in that nectar was originally secreted within the staminal tube; but why has not the one stamen long since cohered? The great difference in structure for fertilisation within the same genus makes one believe that all such points are vary variable. (715/3. Coronilla emerus is of the ordinary papilionaceous type.) With respect to the non-coherence of the one stamen, do examine some flower-buds at a very early age; for parts which are largely developed are often developed to an unusual degree at a very early age, and it seems to me quite possible that the base of the vexillum (to which the single stamen adhered) might thus be developed, and thus keep it separate for a time from the other stamens. The cohering stamens to the right and left of the single one seem to me to be pushed out a little laterally. When you have finished your observations, you really ought to send an account with a diagram to "Nature," recalling your generalisation about the diadelphous structure, and now explaining the exception of Coronilla. (715/4. The observations were published in "Nature," Volume X., 1874, page 169.)
Do add a remark how almost every detail of structure has a meaning where a flower is well examined.
Your observations pleased me so much that I could not sit still for half an hour.
Please to thank Mr. Payne (715/5. Lord Farrer's gardener.) for his remarks, which are of value to me, with reference to Mimosa. I am very much in doubt whether opening the sashes can act by favouring the evaporation of the drops; may not the movement of the leaves shake off the drops, or change their places? If Mr. Payne remembers any plant which is easily injured by drops, I wish he would put a drop or two on a leaf on a bright day, and cover the plant with a clean bell-glass, and do the same for another plant, but without a bell-glass over it, and observe the effects.
Thank you much for wishing to see us again at Abinger, and it is very doubtful whether it will be Coronilla, Mr. Payne, the new garden, the children, E. {Lady Farrer}, or yourself which will give me the most pleasure to see again.
P.S. 1.—It will be curious to note in how many years the rough ground becomes quite uniform in its flora.
P.S. 2.—One may feel sure that periodically nectar was secreted within the flower and then secreted by the calyx, as in some species of Iris and orchids. This latter being taken advantage of in Coronilla would allow of the secretion within the flower ceasing, and as this change was going on in the two secretions, all the parts of the flower would become modified and correlated.
LETTER 716. TO J. BURDON SANDERSON. Down, Tuesday, September 9th {1873}.
(716/1. Sir J. Burdon Sanderson showed that in Dionoea movement is accompanied by electric disturbances closely analogous to those occurring in muscle (see "Nature," 1874, pages 105, 127; "Proc. R. Soc." XXI., and "Phil. Trans." Volume CLXXIII., 1883, where the results are finally discussed).)
I will send up early to-morrow two plants {of Dionoea} with five goodish leaves, which you will know by their being tied to sticks. Please remember that the slightest touch, even by a hair, of the three filaments on each lobe makes the leaf close, and it will not open for twenty-four hours. You had better put 1/4 in. of water into the saucers of the pots. The plants have been kept too cool in order to retard them. You had better keep them rather warm (i.e. temperature of warm greenhouse) for a day, and in a good light.