In this figure, we see that the plane transverse partition, whatever fraction of the cube it cut off, is always of the same dimensions, that is to say is always equal to a2 , or = 1. If one-half of the cube have to be cut off, this plane transverse partition is much the best, for we see by the diagram that a cylindrical partition cutting off an equal volume would have an area about 25%, and a spherical partition would have an area about 50% greater. The point A in the diagram corresponds to the point where the cylindrical partition would begin to have an advantage over the plane, that is to say (as we have seen) when the fraction to be cut off is about one-third, or ·318 of the whole. In like manner, at B the spherical octant begins to have an advantage over the plane; and it is not till we reach the point C that the spherical octant becomes of less area than the quarter-cylinder.

Fig. 139.

The case we have dealt with is of little practical importance to the biologist, because the cases in which a cubical, or rectangular, cell divides unequally, and unsymmetrically, are apparently few; but we can find, as Berthold pointed out, a few examples, for instance in the hairs within the reproductive “conceptacles” of certain Fuci (Sphacelaria, etc., Fig. [139]), or in the “paraphyses” of mosses (Fig. [142]). But it is of great theoretical importance: as serving to introduce us to a large class of cases, in which the shape and the relative dimensions of the original cavity lead, according to the principle of minimal areas, to cell-division in very definite and sometimes unexpected ways. It is not easy, nor indeed possible, to give a generalised account of these cases, for the limiting conditions are somewhat complex, and the math­e­mat­i­cal treatment soon becomes difficult. But it is easy to comprehend a few simple cases, which of themselves will carry us a good long way; and which will go far to convince the student that, in other cases {352} which we cannot fully master, the same guiding principle is at the root of the matter.


The bisection of a solid (or the subdivision of its volume in other definite proportions) soon leads us into a geometry which, if not necessarily difficult, is apt to be unfamiliar; but in such problems we can go a long way, and often far enough for our particular purpose, if we merely consider the plane geometry of a side or section of our figure. For instance, in the case of the cube which we have been just considering, and in the case of the plane and cylindrical partitions by which it has been divided, it is obvious that, since these two partitions extend symmetrically from top to bottom of our cube, that we need only consider (so far as they are concerned) the manner in which they subdivide the base of the cube. The whole problem of the solid, up to a certain point, is contained in our plane diagram of Fig. [138]. And when our particular solid is a solid of revolution, then it is obvious that a study of its plane of symmetry (that is to say any plane passing through its axis of rotation) gives us the solution of the whole problem. The right cone is a case in point, for here the in­ves­ti­ga­tion of its modes of symmetrical subdivision is completely met by an examination of the isosceles triangle which constitutes its plane of symmetry.

The bisection of an isosceles triangle by a line which shall be the shortest possible is a very easy problem. Let ABC be such a triangle of which A is the apex; it may be shewn that, for its shortest line of bisection, we are limited to three cases: viz. to a vertical line AD, bisecting the angle at A and the side BC; to a transverse line parallel to the base BC; or to an oblique line parallel to AB or to AC. The respective magnitudes, or lengths, of these partition lines follow at once from the magnitudes of the angles of our triangle. For we know, to begin with, since the areas of similar figures vary as the squares of their linear dimensions, that, in order to bisect the area, a line parallel to one side of our triangle must always have a length equal to 1 ⁄ √2 of that side. If then, we take our base, BC, in all cases of a length = 2, the transverse partition drawn parallel to it will always have a length equal to 2 ⁄ √2, or = √2. The vertical {353} partition, AD, since BD = 1, will always equal tan β (β being the angle ABC). And the oblique partition, GH, being equal to AB ⁄ √2 = 1 ⁄ (√2 cos β). If then we call our vertical, transverse

Fig. 140.

and oblique partitions, V, T, and O, we have V = tan β; T = √2; and O = 1 ⁄ (√2 cos β), or