V : T : O = tan β ⁄ √2 : 1 : 1 ⁄ (2 cos β).
And, working out these equations for various values of β, we very soon see that the vertical partition (V) is the least of the three until β = 45°, at which limit V and O are each equal to 1 ⁄ √2 = ·707; and that again, when β = 60°, O and T are each = 1, after which T (whose value always = 1) is the shortest of the three partitions. And, as we have seen, these results are at once applicable, not only to the case of the plane triangle, but also to that of the conical cell.
Fig. 141.
In like manner, if we have a spheroidal body, less than a hemisphere, such for instance as a low, watch-glass shaped cell (Fig. [141], a), it is obvious that the smallest possible partition by which we can divide it into two equal halves {354} is (as in our flattened disc) a median vertical one. And likewise, the hemisphere itself can be bisected by no smaller partition meeting the walls at right angles than that median one which divides it into two similar quadrants of a sphere. But if we produce our hemisphere into a more elevated, conical body, or into a cylinder with spherical cap, it is obvious that there comes a point where a transverse, horizontal partition will bisect the figure with less area of partition-wall than a median vertical one (c). And furthermore, there will be an intermediate region, a region where height and base have their relative dimensions nearly equal (as in b), where an oblique partition will be better than either the vertical or the transverse, though here the analogy of our triangle does not suffice to give us the precise limiting values. We need not examine these limitations in detail, but we must look at the curvatures which accompany the several conditions. We have seen that a film tends to set itself at equal angles to the surface which it meets, and therefore, when that surface is a solid, to meet it (or its tangent if it be a curved surface) at right angles. Our vertical partition is, therefore, everywhere normal to the original cell-walls, and constitutes a plane surface.
But in the taller, conical cell with transverse partition, the latter still meets the opposite sides of the cell at right angles, and it follows that it must itself be curved; moreover, since the tension, and therefore the curvature, of the partition is everywhere uniform, it follows that its curved surface must be a portion of a sphere, concave towards the apex of the original, now divided, cell. In the intermediate case, where we have an oblique partition, meeting both the base and the curved sides of the mother-cell, the contact must still be everywhere at right angles: provided we continue to suppose that the walls of the mother-cell (like those of our diagrammatic cube) have become practically rigid before the partition appears, and are therefore not affected and deformed by the tension of the latter. In such a case, and especially when the cell is elliptical in cross-section, or is still more complicated in form, it is evident that the partition, in adapting itself to circumstances and in maintaining itself as a surface of minimal area subject to all the conditions of the case, may have to assume a complex curvature. {355}
Fig. 142.
-shaped partitions: A, from Taonia atomaria (after Reinke); B, from paraphyses of Fucus; C, from rhizoids of Moss; D, from paraphyses of Polytrichum.