This seems to be the origin of the compound forms of mules, which evidently partake of both parents, but principally of the male parent. In this production of chimeras the antients seem to have indulged their fancies, whence the sphinxes, griffins, dragons, centaurs, and minotaurs, which are vanished from modern credulity.

It would seem, that in these unnatural conjunctions, when the nutriment deposited by the female was so ill adapted to stimulate the living filament derived from the male into action, and to be received; or embraced by it, and combined with it into organization, as not to produce the organs necessary to life, as the brain, or heart, or stomach, that no mule was produced. Where all the parts necessary to life in these compound animals were formed sufficiently perfect, except the parts of generation, those animals were produced which are now called mules.

The formation of the organs of sexual generation, in contradistinction to that by lateral buds, in vegetables, and in some animals, as the polypus, the tænia, and the volvox, seems the chef d'œuvre, the master-piece of nature; as appears from many flying insects, as in moths and butterflies, who seem to undergo a general change of their forms solely for the purpose of sexual reproduction, and in all other animals this organ is not complete till the maturity of the creature. Whence it happens that, in the copulation of animals of different species, the parts necessary to life are frequently completely formed; but those for the purpose of generation are defective, as requiring a nicer organization; or more exact coincidence of the particles of nutriment to the irritabilities or appetencies of the original living filament. Whereas those mules, where all the parts could be perfectly formed, may have been produced in early periods of time, and may have added to the numbers of our various species of animals, as before observed.

As this production of mules is a constant effect from the conjunction of different species of animals, those between the horse and the female ass always resembling the horse more than the ass; and those, on the contrary, between the male ass and the mare, always resembling the ass more than the mare; it cannot be ascribed to the imagination of the male animal which cannot be supposed to operate so uniformly; but to the form of the first nutritive particles, and to their peculiar stimulus exciting the living filament to select and combine them with itself. There is a similar uniformity of effect in respect to the colour of the progeny produced between a white man, and a black woman, which, if I am well informed, is always of the mulatto kind, or a mixture of the two; which may perhaps be imputed to the peculiar form of the particles of nutriment supplied to the embryon by the mother at the early period of its existence, and their peculiar stimulus; as this effect, like that of the mule progeny above treated of, is uniform and consistent, and cannot therefore be ascribed to the imagination of either of the parents.

Dr. Thunberg observes, in his Journey to the Cape of Good Hope, that there are some families, which have descended from blacks in the female line for three generations. The first generation proceeding from an European, who married a tawny slave, remains tawny, but approaches to a white complexion; but the children of the third generation, mixed with Europeans, become quite white, and are often remarkably beautiful. V. i. p. 112.

When the embryon has produced a placenta, and furnished itself with vessels for selection of nutritious particles, and for oxygenation of them, no great change in its form or colour is likely to be produced by the particles of sustenance it now takes from the fluid, in which it is immersed; because it has now acquired organs to alter or new combine them. Hence it continues to grow, whether this fluid, in which it swims, be formed by the uterus or by any other cavity of the body, as in extra-uterine gestation; and which would seem to be produced by the stimulus of the fetus on the sides of the cavity, where it is found, as mentioned before. And thirdly, there is still less reason to expect any unnatural change to happen to the child after its birth from the difference of the milk it now takes; because it has acquired a stomach, and lungs, and glands, of sufficient power to decompose and recombine the milk; and thus to prepare from it the various kinds of nutritious particles, which the appetencies of the various fibrils or nerves may require.

From all this reasoning I would conclude, that though the imagination of the female may be supposed to affect the embryon by producing a difference in its early nutriment; yet that no such power can affect it after it has obtained a placenta, and other organs; which may select or change the food, which is presented to it either in the liquor amnii, or in the milk. Now as the eggs in pullets, like the seeds in vegetables, are produced gradually, long before they are impregnated, it does not appear how any sudden effect of imagination of the mother at the time of impregnation can produce any considerable change in the nutriment already thus laid up for the expected or desired embryon. And that hence any changes of the embryon, except those uniform ones in the production of mules and mulattoes, more probably depend on the imagination of the male parent. At the same time it seems manifest, that those monstrous births, which consist in some deficiencies only, or some redundancies of parts, originate from the deficiency or redundance of the first nutriment prepared in the ovary, or in the part of the egg immediately surrounding the cicatricula, as described above; and which continues some time to excite the first living filament into action, after the simple animal is completed; or ceases to excite it, before the complete form is accomplished. The former of these circumstances is evinced by the eggs with double yolks, which frequently happen to our domesticated poultry, and which, I believe, are so formed before impregnation, but which would be well worth attending to, both before and after impregnation; as it is probable, something valuable on this subject might be learnt from them. The latter circumstance, or that of deficiency of original nutriment, may be deduced from reverse analogy.

There are, however, other kinds of monstrous births, which neither depend on deficiency of parts, or supernumerary ones; nor are owing to the conjunction of animals of different species; but which appear to be new conformations, or new dispositions of parts in respect to each other, and which, like the variation of colours and forms of our domesticated animals, and probably the sexual parts of all animals, may depend on the imagination of the male parent, which we now come to consider.

[VI]. [1]. The nice actions of the extremities of our various glands are exhibited in their various productions, which are believed to be made by the gland, and not previously to exist as such in the blood.

Thus the glands, which constitute the liver, make bile; those of the stomach make gastric acid; those beneath the jaw, saliva; those of the ears, ear-wax; and the like. Every kind of gland must possess a peculiar irritability, and probably a sensibility, at the early state of its existence; and must be furnished with a nerve of sense, or of motion, to perceive, and to select, and to combine the particles, which compose the fluid it secretes. And this nerve of sense which perceives the different articles which compose the blood, must at least be conceived to be as fine and subtile an organ, as the optic or auditory nerve, which perceive light or sound. See Sect. [XIV. 9].