But it appears from dissection, that the tympanum is not the immediate organ of hearing, but that like the humours and cornea of the eye, it is only of use to prepare the object for the immediate organ. For the portio mollis of the auditory nerve is not spread upon the tympanum, but upon the vestibulum, and cochlea, and semicircular canals of the ear; while between the tympanum and the expansion of the auditory nerve the cavity is said by Dr. Cotunnus and Dr. Meckel to be filled with water; as they had frequently observed by freezing the heads of dead animals before they dissected them; and water being a more dense fluid than air is much better adapted to the propagation of vibrations. We may add, that even the external opening of the ear is not absolutely necessary for the perception of sound: for some people, who from these defects would have been completely deaf, have distinguished acute or grave sounds by the tremours of a stick held between their teeth propagated along the bones of the head, (Haller. Phys. T. V. p. 295).
Hence it appears, that the immediate organ of hearing is not affected by the particles of the air themselves, but is stimulated into animal motion by the vibrations of them. And it is probable from the loose bones, which are found in the heads of some fishes, that the vibrations of water are sensible to the inhabitants of that element by a similar organ.
The motions of the atmosphere, which we become acquainted with by the sense of touch, are combined with its solidity, weight, or vis intertiæ; whereas those, that are perceived by this organ, depend alone on its elasticity. But though the vibration of the air is the immediate object of the sense of hearing, yet the ideas, we receive by this sense, like those received from light, are only as a language, which by acquired associations acquaints us with those motions of tangible bodies, which depend on their elasticity; and which we had before learned by our sense of touch.
[V]. Of Smell and of Taste.
The objects of smell are dissolved in the fluid atmosphere, and those of taste in the saliva, or other aqueous fluid, for the better diffusing them on their respective organs, which seem to be stimulated into animal motion perhaps by the chemical affinities of these particles, which constitute the sapidity and odorosity of bodies with the nerves of sense, which perceive them.
Mr. Volta has lately observed a curious circumstance relative to our sense of taste. If a bit of clean lead and a bit of clean silver be separately applied to the tongue and palate no taste is perceived; but by applying them in contact in respect to the parts out of the mouth, and nearly so in respect to the parts, which are immediately applied to the tongue and palate, a saline or acidulous taste is perceived, as of a fluid like a stream of electricity passing from one of them to the other. This new application of the sense of taste deserves further investigation, as it may acquaint us with new properties of matter.
From the experiments above mentioned of Galvani, Volta, Fowler, and others, it appears, that a plate of zinc and a plate of silver have greater effect than lead and silver. If one edge of a plate of silver about the size of half a crown-piece be placed upon the tongue, and one edge of a plate of zinc about the same size beneath the tongue, and if their opposite edges are then brought into contact before the point of the tongue, a taste is perceived at the moment of their coming into contact; secondly, if one of the above plates be put between the upper lip and the gum of the fore-teeth, and the other be placed under the tongue, and their exterior edges be then brought into contact in a darkish room, a flash of light is perceived in the eyes.
These effects I imagine only shew the sensibility of our nerves of sense to very small quantities of the electric fluid, as it passes through them; for I suppose these sensations are occasioned by slight electric shocks produced in the following manner. By the experiments published by Mr. Bennet, with his ingenious doubler of electricity, which is the greatest discovery made in that science since the coated jar, and the eduction of lightning from the skies, it appears that zinc was always found minus, and silver was always found plus, when both of them were in their separate state. Hence, when they are placed in the manner above described, as soon as their exterior edges come nearly into contact, so near as to have an extremely thin plate of air between them, that plate of air becomes charged in the same manner as a plate of coated glass; and is at the same instant discharged through the nerves of taste or of sight, and gives the sensations, as above described, of light or of saporocity; and only shews the great sensibility of these organs of sense to the stimulus of the electric fluid in suddenly passing through them.
[VI]. Of the Sense of Heat.
There are many experiments in chemical writers, that evince the existence of heat as a fluid element, which covers and pervades all bodies, and is attracted by the solutions of some of them, and is detruded from the combination of others. Thus from the combinations of metals with acids, and from those combinations of animal fluids, which are termed secretions, this fluid matter of heat is given out amongst the neighbouring bodies; and in the solutions of salts in water, or of water in air, it is absorbed from the bodies, that surround them; whilst in its facility in passing through metallic bodies, and its difficulty in pervading resins and glass, it resembles the properties of the electric aura; and is like that excited by friction, and seems like that to gravitate amongst other bodies in its uncombined state, and to find its equilibrium.