Physiology and Hygiene for Secondary Schools

by Francis M. Walters

Fig. 153—Diagram showing the divisions of cochlear canal.

The Cochlea is the part of the internal ear directly concerned in hearing. It consists of a coiled tube which makes two and one half turns around a central axis and bears a close resemblance to a snail shell (Figs. 151 and 152). It differs in plan from a snail shell, however, in that its interior space is divided into three distinct channels, or canals. These lie side by side and are named, from their relations to other parts, the scala vestibula, the scala tympani, and the scala media. Any vertical section of the cochlea shows all three of these channels (Fig. 153).

[pg 363]The Scala Vestibula and the Scala Tympani appear in cross section as the larger of the canals. The former, so named from its connection with the vestibule, occupies the upper position in all parts of the coil. The latter lies below at all places, and is separated from the channels above partly by a margin of bone and partly by a membrane. It receives its name from its termination at the tympanum, or middle ear, from which it is separated only by a thin membrane.[122] Both the scala vestibula and the scala tympani belong to the outer portion of the internal ear and are, for this reason, filled with the perilymph. At their upper ends they communicate with each other by a small opening, making by this means one continuous canal through the cochlea. This canal passes from the vestibule to the tympanum and, in so doing, goes entirely around

The Scala Media.—This division of the cochlea lies parallel to and between the other two divisions. It is above the scala tympani and below the scala vestibula, and is separated from each by a membrane. The scala media belongs to the membranous portion of the internal ear and is, therefore, filled with the endolymph. It receives the terminations of fibers from the auditory nerve and may be regarded as the true sense organ of hearing. The nerve fibers terminate upon the membrane known as the basilar membrane, which separates it from the scala tympani. This membrane extends the length of the cochlear canals, and is stretched between a projecting shelf of bone on one side and the outer wall of the cochlea on the other. It is covered with a layer of epithelial cells, some of which have small, hair-like projections and are known as the hair cells. Above the membrane, and resting partly upon it, are two[pg 364] rows of rod-like bodies, called the rods of Corti. These, by leaning toward each other, form a kind of tunnel beneath. They are exceedingly numerous, numbering more than 6000, and form a continuous series along the margin of the membrane.

Fig. 154—Diagram illustrating passage of sound waves through the ear.

How We Hear.—The sound waves which originate in vibrating bodies are transmitted by the air to the external ear. Passing through the auditory canal, the waves strike against the membrana tympani, setting it into vibration. By the bridge of bones and the air within the middle ear the vibrations are carried to and concentrated upon the liquid in the internal ear (Fig. 154). From here the vibrations pass through the channels of the cochlea and set into vibration the contents of the scala media and different portions of the basilar membrane. This serves as a stimulus to the fibers of the auditory nerve, causing them to transmit impulses which, on passing to the brain, produce the sensation of hearing.

Much of the peculiar structure of the cochlea is not understood. Its minute size and its location in the temporal bone make its study extremely difficult. The connection of the scala vestibula with the scala tympani, and this with the middle ear, is necessary for the passage of vibrations through the internal ear. Its liquids, being practically incompressible and surrounded on all sides by bones, could not otherwise yield to the movements of the stapes. (See Practical Work.) The rods of Corti are thought to act as dampers on the basilar membrane, to prevent the continuance of vibrations when once they are started.

Detection of Pitch.—The method of detecting tones of different pitch[pg 365] is not understood. Several theories have been advanced with reference to its explanation, one of the most interesting being that proposed by Helmholtz. This theory is based on our knowledge of sympathetic vibrations. The basilar membrane, while continuous throughout, may be regarded as made up of many separate cords of different lengths stretched side by side. A tone of a given pitch will set into vibration only certain of these cords, while tones of different pitch will set others into vibration.