B. arnouxi, from the seas of New Zealand, is the only species of this genus which is well known. It is 30 or 32 feet in length, and is of a velvety black colour, with a greyish belly. Instead of lowing like a cow, this Whale has been described as "bellowing like a bull"! A singular and somewhat inexplicable fact has been stated of this species. The teeth were said to be protrusible, and Sir James Hector stated that the teeth were imbedded "in a tough cartilaginous sac which adheres loosely in the socket of the jaw, and is moved by a series of muscular bundles that elevate or depress it." Sir William Flower justly observed that these statements "accord so little with anything hitherto known in mammalian anatomy that further observations on the subject are extremely desirable." Like other Ziphioids, Berardius feeds mainly, if not entirely, upon cuttle-fish, a prey eminently suited to their almost toothless mouths. It is not known whether Berardius has the
Ziphioid grooves upon the throat. Nothing is known of the structure of the internal viscera of this Whale. It appears not to be really limited to the region of New Zealand, as is often stated, for Malm has lately described a skull (Berardius vegae) from Bering's Straits.[[241]]
Mesoplodon[[242]] is a world-wide genus embracing a number of species; on the lowest estimate seven species can be distinguished, and Sir W. Flower would add two more. These are moderate-sized Whales, 15 to 17 feet in length. In the skull the mesethmoid is ossified; the nasals are sunk between the upper ends of the premaxillae. There are but a single pair of teeth in the mandible attached to nearly the middle of its length (whence the generic name). The vertebral formula is C 7, D 9 or 10, L 10 or 11, Ca 19 or 20. The sternum consists of four or five pieces. The amount to which the cervical vertebrae are fused varies; but some are always fused.
The only species which has ever been stranded on the shores of this country is M. bidens, an example of which was described many years ago as the "Toothless Whale of Havre"; it was an old animal which had probably lost its teeth. Nevertheless it received the separate generic and specific name of Aodon dalei. The animal lived for two days out of the water, and made a sound like the "lowing of a cow." An instance of the rarity of the Whales of this genus is afforded by M. europaeus, of which only a single skull is known; this was extracted from a dead body, found floating, about the year 1840. It has never appeared since. M. layardi is remarkable on account of the very large size of its strap-shaped teeth; these curve over the upper jaw in such a way as to prevent the animal from fully opening its jaws. The case is curiously paralleled by the Sabre-toothed Tiger. This species is antarctic in range. From the opposite extremity of the globe comes M. stejnegeri, again known by but a single skull. It is singular on account of the large size of the brain case, and is a native of Bering's Straits. M. hectori has its two teeth situated quite at the extremity of the mandible, and in this feature approximates to the genus Berardius. It was, indeed, confounded with that genus by one naturalist.
Ziphius is a genus which is also of world-wide range. Here again the number of species is at present merely a matter of opinion. The prevalent impression, however, is that but a single species exists, which will therefore have the name of Z. cavirostris. The genus (and for the matter of that the species too) may be thus characterised in comparison with its allies. The mesethmoid is ossified as in Mesoplodon, but the nasals joined together form the vertex of the skull. There are two teeth near the symphysis of the mandible, besides the usual small and "functionless" teeth in the upper jaw. The vertebral formula is C 7, D 9 or 10, L 11, Ca 21.
The throat of a Ziphius from New Zealand was described by Messrs. Scott and Parker[[243]] as having three grooves on each side. Whether this form is the same as von Haast's Z. novae zelandiae is a matter of doubt; but the individual to which his name has been applied was 26 feet long, and had but a single groove on each side. Even in the external characters of many Whales many points require clearing up. Our knowledge of Ziphius dates from the year 1804, when a skull "completely petrified in appearance" was picked up upon the Mediterranean coast of France, and described by the great Cuvier. It was forty years before another specimen was found. In the New Zealand specimen of von Haast already referred to, the body was scored by numerous lacerations. These wounds may have been due to fights among the Whales themselves; the forwardly-situated teeth would be capable of inflicting such wounds. But it has also been stated that the armed suckers of gigantic cuttle-fish are responsible for these scratches.
Hyperoodon is the most easily-distinguishable genus of Ziphioid Whales. Its characters are the following:—The skull has enormously-developed maxillary crests in the adult male; the mesethmoid is not fully ossified. There is but a single tooth to each ramus of the lower jaw, besides, of course, the usual small teeth in the upper jaw. The vertebral formula is C 7, D 9, L 9, Ca 18. The cervicals are fused into one mass, more or fewer being free in other Ziphioids. The sternum consists of three pieces only, the last of which is bifid posteriorly.
The name Hyperoodon was given to this Whale by Colonel Lacepède on account of the rough papillae upon the palate, which
were mistaken by that observer for teeth. It is curious that the name is really appropriate in spite of this mistake, though of course it would be so to all the Ziphioids. In more than one feature this genus comes nearest of all the Ziphiinae to Physeter. Its enormous maxillary crests are paralleled in that Whale; but in Hyperoodon their great thickness contrasts with the thinness of those of the Cachalot. The correspondence in the attachment of a rib to its vertebra by both heads is noteworthy. It is remarkable that in this particular Hyperoodon is more like Physeter than the supposed nearest ally of the latter—Kogia.