Of this genus two species are known. The best known is the common northern H. rostratum (with many aliases); the second species from the southern hemisphere, H. planifrons, is only known from a single water- and pebble-worn skull. Its identification, however, depends upon the known accuracy of the late Sir William Flower.
The northern species (Hyperoodon rostratum) has often been recorded upon our own coasts; the first record of the stranding of this Whale was in the year 1717. In that year an example was found at Maldon, in Essex. Like the Beluga, Hyperoodon rostratum gets lighter in colour with advancing years. The young are black; the old animals pale brown with some white about them. The under surface, however, is always greyish white. The length of this Whale reaches to at any rate 30 feet. But John Hunter had a specimen which he believed to be 40 feet in length. The specimen, however, consisted only of a skull, so that error might have crept in. It has already been mentioned that the old males have enormous maxillary crests. According to M. Bouvier, who has lately made an exhaustive examination of the anatomy of this Whale,[[244]] the females occasionally exhibit the same crests, which are thus presumably of the nature of spurs sometimes seen in old females among the Gallinaceous birds. The number of grooves upon the throat is in dispute in this Whale as in Ziphius. One pair is the usual allowance; but Kükenthal found four in some embryos studied by him. Attention has already been called to the voice of Ziphioid Whales. Hyperoodon neither "lows" nor "bellows," but "sobs"! Hyperoodon rostratum is a gregarious Whale, going about in herds, or "gams" as they should technically be termed, of four to ten or even fifteen. This Whale
can leap right out of the water, and while in the air can turn its head from side to side, a capability which has not been mentioned in any other Whale. It can also stay under water for an unusually long period. Captain Gray,[[245]] who has made an accurate study of this species, states that so long a period as two hours is the limit of endurance; this event occurred in the case of a harpooned Whale.
Fam. 2. Delphinidae.—This family, which includes the greater number of Cetacea, may thus be characterised:—Whales of small to moderate size. Teeth as a rule numerous, and present in the upper as well as in the lower jaw. Maxillae without large crests; the pterygoids, often meeting in the middle line, enclose an air space open behind. The anterior (five to eight) ribs are two headed, the posterior with tubercular head only. The sternal ribs are ossified.
The Dolphins and Porpoises, as already stated, embrace the greater number of existing species of Whales. Sir W. Flower and others who have followed him, allow nineteen genera. But as to the exact number of known species there is much uncertainty. That very careful observer, Mr. True, considers[[246]] that there are fifty which demand recognition. As many as one hundred have received names. The matter is one which is perhaps barely ripe for decision. All the Dolphin tribe are, for Whales, smallish animals. The Killer Whale, Orca, is the only genus (or species?) which usually attains to more than moderate bulk. The rather mysterious Delphinus coronatus, 36 feet in length, of M. de Fréminville, would seem to be a Ziphioid; it was described as having a very pointed beak, and as having the dorsal fin situated near the tail; such characters suggest a Mesoplodon.
The genus Delphinapterus, the Beluga or White Whale, consists of but a single species, though as usual more than one name has been given to supposed different species. It is characterised as a genus by the following assemblage of structural features:—It has only eight to ten teeth occupying the anterior part of the jaws only. All the cervical vertebrae are free and unjoined. The vertebral formula is C 7, D 11 (or 12), L 9, Ca 23. The pterygoids are wide apart, though they converge as if about to meet at their posterior ends. There is no dorsal fin. The colour is white.
The Beluga is a northern species purely. The reputed form, D. kingii, was said to come from Australian seas; but there seems to have been an error in this statement. It is interesting to note that the white colour, so characteristic of the genus and species, is not found in the young, which are blackish. They gradually pale as they advance towards maturity. Delphinapterus leucas reaches a length of 10 feet, and like other Porpoises will ascend rivers in search of food. It is said to be specially addicted to salmon. Among the contents of the stomach have been found quantities of sand. But this habit of swallowing sand or pebbles has been noted in other Whales. Whether it is or is not accidental (taken in with ground-living food), it seems hardly likely that it is used for purposes of ballast! The Beluga has a voice; but the name "Sea Canary" is hardly suitable to it. A specimen of this species, recently described from the shores of Scotland (it is often thrown up upon our coasts), which had got entangled in the stakes of a new net, was regarded by the natives, on account of its white colour, as a ghost. Externally, besides its colour, the Beluga is remarkable for possessing a distinct neck, which is correlated of course with the freedom of the cervical vertebrae, and is also seen in Platanistidae.
The Narwhal (Monodon) is closely allied in structure to the last genus. It has the following anatomical characters:—The teeth are reduced to a single "horn" in the upper jaw, which is rudimentary in the female. The neck vertebrae are free. The vertebral formula is C 7, D 11, L 6, Ca 26. The pterygoids are as in Delphinapterus, and, as in that genus, there are no hairs upon the face or dorsal fin.
This genus is of course most obviously characterised by the twisted tusk of the male, which is occasionally double. This tusk has given to the only species of the genus, M. monoceros, both its generic and specific name. The animal has a spotted colour; but, as in the case of the Beluga, old animals tend to become white. The use of its horn to Monodon has been debated. In the first place it is clearly a secondary sexual character. The males have been observed to cross their horns like rapiers in a fencing match. It may be that they are used in more serious combats. An ingenious suggestion is that the long and strong tusk enables its possessor to break the thick ice and