The genus Tursio must be carefully distinguished from
Tursiops. It has no dorsal fin, the teeth are small and numerous (forty-four), and the pterygoids are separate. There are two species, T. borealis and T. peronii, the former being northern and the latter more widely spread.
The genus Cephalorhynchus has for its chief characters the following:—Teeth twenty-five to thirty-one, small and sharp. Pterygoids widely separated. Dorsal fin not falcate, but triangular or ovate in form. Beak not well marked off from the head. The species of this genus are all southern in range; four are perhaps to be allowed.
Fam. 3. Platanistidae.—This family of Odontocetes may be distinguished from the Dolphins by the following assemblage of structural features:—Cervical vertebrae all free, and each one of some length (for a Cetacean). Jaws long and narrow, with a considerable length of symphysis. Teeth very numerous.
This very meagre series of differential characters is largely due to Pontoporia on the Platanistid side, and to Monodon and Delphinapterus upon the Delphinid side. Otherwise the family Platanistidae would be extremely distinct. The two last-named genera have separate cervical vertebrae, and in the Beluga at any rate this is expressed externally by a quite distinct neck. Moreover, as Mr. True has pointed out, the pterygoid bones have not the involuted cavity below which characterises other Dolphins; and they have, what other Dolphins have not, an articulation outwards with the roofing bones of the skull. Sir W. Flower described the fact that in Inia (and the same occurs in Pontoporia) the palatines are separated from each other by the intervention of the vomer. In this feature they resemble certain Ziphioids, Berardius, Oulodon (= Mesoplodon) grayi, and Hyperoodon. The true Dolphins also appear to show the same intervention of the vomer in a few cases. There is nothing, therefore, distinctive from the Delphinidae in this feature.
The existence of cartilaginous sternal ribs in Inia and Platanista shows affinity between these two genera and the Physeteridae. Pontoporia is Dolphin-like in this particular, as it is also in the mode of articulation of the ribs with the vertebral column. But this last matter has already been dealt with. The principal reason for placing Pontoporia with the other two genera is the close resemblance which its skull bears to that of Inia.
The first genus of this family which will be noticed is Platanista.
The following are its main characters:—Dorsal fin absent. Eyes rudimentary. Pectoral fins large and truncated at the extremity. Teeth, about twenty-nine in each half of each jaw. Scapula with the acromion coinciding with its anterior edge. Skull with enormous maxillary crests, and with the palatines entirely concealed by the pterygoids. The length of the above definition will serve to indicate how anomalous in many particulars is the structure of this "Dolphin."
There is apparently but one species, P. gangetica, the "Susu." The Indian vernacular name is derived from the sound that the animal makes when spouting. It is an inhabitant of the Ganges and the Indus, together with their tributaries, and ascends very high up its streams. It is also thought to be purely fluviatile and never to desert the rivers for the sea. Platanista lives chiefly by grubbing in the mud for prawns and fish. Grains of rice have also been found in the stomach, but this would seem to be accidental. The long snout of the Susu has been compared to the long snout of the Gharial, a native of the same region. This Whale grows to a length of over 9 feet, but this length is exceptional. Its anatomy has been elaborately described by Dr. Anderson.[[252]]
The next genus, Inia, is thus to be characterised:—Dorsal fin rudimentary; pectorals large and ovate. Teeth, as many as thirty-two on each side, often with an additional tubercle. Skull without large maxillary crests; palatines not hidden by pterygoids, but divided by vomer. The vertebrae of this genus are few in number, only forty-one in all, which are thus distributed: C 7, D 13, L 3, Ca 18. The peculiarities of the vertebral column are several. In the first place, as has been mentioned in the definition of the family, all the cervicals are separate and individually of some length. Secondly, the axis has a better trace of an odontoid process than in any other Whale except Platanista, where it is even more obvious. The lumbar region is remarkable on account of its restriction to three vertebrae. The sternum, by what we must regard as convergence, is somewhat like that of the Whalebone whales. It consists of one piece only, of a roughly-oval form, to which apparently only two pairs of (cartilaginous) sternal ribs are attached. In the fore-limb the proportions between the humerus and the radius are more like