those of terrestrial mammals; i.e. the humerus is distinctly the longer, the converse usually obtaining among Whales. But Platanista again agrees with Inia. The teeth are remarkable for the fact that the hindermost ones of the series have an additional lobe; they are not purely conical as are those of Whales generally.
There is but one species, Inia geoffrensis, which inhabits the Amazons, and grows to a length of 8 feet. Its colour variations are rather extraordinary, unless they can be set down to sex, which has been denied. Some individuals are wholly pink; others are black above and pink beneath. This Whale is believed by the Indians to attack a man in the water, and it is added that the Sotalia of the same streams will defend him from these attacks! Naturally, therefore, superstitious reverence attaches to this Dolphin, which is tiresome to the naturalist who wants specimens, as Professor Louis Agassiz found.
In the genus Pontoporia[[253]] the dorsal fin is well developed and falcate. The teeth are very numerous, 200 in all. The ribs articulate as in Dolphins. The skull closely resembles that of Inia, and the scapula is, as in that genus, "normal."
The proper name for Pontoporia is really Stenodelphis, which name was first used by Gervais a month or two before Gray, who separated it from the vague Delphinus of its original discoverer, Gervais himself. It has a longer snout than Inia, which, being bent towards the extremity in a downward direction, curiously suggests the skull of a Curlew. In details, however, the skull is exceedingly like that of Inia. It is nearly symmetrical. The vertebral formula appears to be the following:—C 7, D 10, L 5, Ca 20 = 42, just one over the number of the vertebrae in Inia. The sternum is in two pieces. Of the ten pairs of ribs the first three are double-headed. These and the next have sternal moieties joining the sternum, of which the first three are ossified, the last being apparently merely a ligament.
There is a single species of the genus, P. blainvillii. This Whale is described by Mr. Lydekker as being of a clear brown colour, harmonising with the waters of the estuary of the Amazons and the La Plata which it inhabits. The same colour characterises Sotalia pallida of those parts of the world, and
may be a colour adaptation. But the extant accounts of the colour of this Dolphin vary—quite possibly in accordance with real variations, such as are exhibited by Inia already spoken of. Pontoporia blainvillii is a smallish Dolphin some 4 feet in length.
Fossil Odontocetes.—Several of the existing genera of Dolphins are also known in a fossil condition, as well as Ziphioid Whales closely related to existing forms. We shall deal here only with a few genera of fossil Odontocetes which depart in their structure from existing forms.
The genus Physodon is Miocene, and has been found in Patagonia. It appears to be most nearly allied to the Physeteridae, but should probably form a distinct family. Physodon was not so large as Physeter, the skull measuring only some 10 feet. It thus comes nearer in point of size to Kogia, and it is interesting to note that its relatively-shorter snout is also suggestive of the dwarf Cachalot. The general outline of the skull is, however, more like that of Physeter, and there is the same deep cavity for the lodgment of spermaceti. The main feature of interest in the skull is the presence of teeth in both jaws, and the fact that two or three are lodged in the premaxillae. This is precisely what is found in the most ancient Whales, the Zeuglodonts.
Extinct Dolphins, apparently referable to the Platanistidae, are the most numerous among the earlier forms of Cetaceans, and it is significant that the earliest known forms of these go back to the Eocene.
The genus Iniopsis of Mr. Lydekker,[[254]] with one species, I. caucasica, comes from rocks which seem to be of that age. The back part of the skull of this animal, the only part of the skull known, has the same squarish excavation of the maxillaries that characterises Inia and Pontoporia. Its lower jaw was slender and possessed numerous teeth.