Fig. 66.—Red Kangaroo. Macropus rufus. × 1⁄18.
Sub-Fam. 1. Macropodinae.—The genus Macropus includes not only the Kangaroos but also the Wallabies, which are really indistinguishable, though they have sometimes been placed in a separate genus Halmaturus. The genus thus enlarged contains twenty-three species. It may be thus characterised: the ears are long, the rhinarium is usually naked, but in M. giganteus and others a band of hairs descends to the upper lip; a naked band extends from the ankle to the pads on the digits, which is interrupted in M. rufus by a band of hairs just in front of the digits. The mammae are four. The tail is not bushy,
but is crested in M. irma. They are for the most part found on the Australian continent, but some species are found in the islands to the north which belong to the Australian region. Thus M. brunii, which is of interest as the first Kangaroo seen by a European, is a native of the Aru islands. A specimen of this animal, which was then living in the garden of the Dutch governor of Batavia, was described by Bruyn in the year 1711. M. rufus, the largest member of the group, is remarkable for the red secretion which adorns the neck of the male. It is caused by particles which have the appearance and colour of carmine. M. giganteus is not, as its specific name might imply, the "giant" of the race; its dimensions are given as 5 feet, while M. rufus is said to attain a length of 5 feet 5 inches, exclusive (in both cases) of the tail.
The account which Sir Joseph Banks gives[[75]] in his diary of the Kangaroo is interesting, since he was one of the first naturalists to see that creature. In July 1770 it was reported to him that an "animal as large as a greyhound, of a mouse colour, and very swift" had been seen by his people. A little later he was surprised to observe that the animal "went only upon two legs, making vast bounds just as the jerboa does." The second lieutenant killed one of these Kangaroos, of which Sir Joseph Banks wrote that "to compare it to any European animal would be impossible, as it has not the least resemblance to any one I have seen. Its fore-limbs are extremely short and of no use to it in walking; its hind, again, as disproportionately long; with these it hops seven or eight feet at a time, in the same manner as the jerboa, to which animal indeed it bears much resemblance, except in size, this being in weight 38 lbs., and the jerboa no larger than a common rat." The beast was killed and eaten, and proved excellent meat. Sir Joseph Banks' observations upon the leaping of the Kangaroo are of interest, because it is often asserted that the tail is largely made use of as a third foot or as a support. Mr. Aflalo declares in the most positive way that after repeatedly examining the tracks upon soft sand immediately after the animal had passed, not the very faintest trace of the impression of the tail could be discovered. The leaps of a big Kangaroo seem to be somewhat greater than is recorded
by Banks. It is said that 15 or even 20 feet are covered at a bound, and in bound after bound. But in walking slowly it can be readily seen from an inspection of Kangaroos at the Zoological Society's Gardens that the animal does rest upon its tail, which with the hind-legs forms a tripod.
Petrogale with six species comes next to Macropus, and is indeed only to be differentiated from it by the thickly-haired and more slender tail, which is not used, as it is sometimes in the Kangaroos, as an extra hind-limb. The Rock-Kangaroos live among rocks, which they climb, and from which they leap; and the tail acts rather as a balancing pole. The most elaborate account of the anatomy of Petrogale known to me is by Mr. Parsons.[[76]] The dentition as given by Mr. Thomas is I 3/1 C 0/0 Pm 2/2 M 4/4—that of Macropus without the occasionally occurring canine of the upper jaw. The osteological characters which separate it from Macropus are quite insignificant. Mr. Parsons mentions a wormian bone, "os epilepticum," at the junction of the coronal and sagittal sutures. It was found to occur in two out of five skulls examined, and appears not to occur in other Kangaroos. The palatine foramina of Petrogale are so large that the posterior part of the bone is only a narrow thickened ridge. The small intestine of P. xanthopus is 102 inches long, the large intestine 44 inches. The caecum has a length of 6 inches, and is not sacculated, differing in this from the caecum of Macropus major. The best known species are P. xanthopus and P. penicillata. The genus is confined to Australia itself, and does not enter Tasmania.
Onychogale includes the so-called "Nail-tailed Wallabies," which have a thorn at the end of the tail, reminding one of the Lion and the Leopard, whose tails have a similar armature. The muffle is hairy. Three species are allowed by Mr. Thomas.
Lagorchestes has, like the last genus, the rhinarium, i.e. that part of the nose immediately surrounding the nostrils, hairy instead of smooth as in the Kangaroos proper. It is distinguished from Onychogale by the absence of the terminal callosity to the tail, which is rather short. The name Hare-Kangaroo is given to the members of this genus (three species) on account of their exceeding fleetness. This genus is limited to Australia itself. L. conspicillatus is said to present "a remarkable
resemblance to the English hare," and L. leporoides was so called by Gould on account of general appearance as well as face.