Dorcopsis has shorter hind-legs than Macropus, and a naked muffle. The ears are small. The structure of D. luctuosa has been studied by Garrod,[[77]] who pointed out the existence of four enlarged hair follicles on the neck near the mandibular symphysis. These are, however, represented in the next genus Dendrolagus, and occur also in Petrogale. The limbs are not so disproportionate as in Macropus, and the tail is naked at the tip.
Dorcopsis and the next genus to be described, Dendrolagus, differ from Macropus and its immediate allies, Petrogale and Lagorchestes, in a number of anatomical points. In the first place, the premolars are twice the size of those of Macropus, and they have a characteristic pattern not observable in the Kangaroos. This consists of a median ridge (the whole tooth being rather prismatic in shape), with lateral ridges at right angles to it. The upper canines are developed, but are minute.
The stomach is not quite like that of Macropus, though built upon a similar plan. The blind cardiac extremity is a single, not a double cul-de-sac; in this it is like that of Petrogale. The distribution of the squamous, white, oesophageal epithelium is very much like that of Dendrolagus. In both genera the orifice of the oesophagus into the stomach is guarded by two strong longitudinal folds, which run for some distance towards the pylorus. In Dendrolagus, at any rate, this tract is bordered on each side by glandular patches. In Dendrolagus, moreover, the squamous epithelium does not extend into the cardiac cul-de-sac. This latter is separated from the rest of the stomach by two slightly diverging folds, which are faintly represented in Petrogale and in Halmaturus. In the last two genera the folds surrounding the oesophageal orifice are but slightly represented; better in Halmaturus than in Petrogale. But there are not the patches of glands already referred to. The small intestine of Dorcopsis is 97 inches in length, the large being 32, i.e. proportionately long, as in Marsupials generally. The small caecum (2½ inches) is not sacculated.
The spleen is Macropodine, being
-shaped or
-shaped. The differences between Dorcopsis and the evidently closely allied Dendrolagus will be further considered under the description of
the latter. Dorcopsis is confined to New Guinea, and contains three species, viz. D. muelleri, D. luctuosa, and D. macleani. D. muelleri has a striking resemblance to Macropus brunii, with which it has been confounded. Though intermediate between Macropus and Dendrolagus, these Kangaroos are not arboreal.
The genus Dendrolagus is remarkable for its un-kangaroo-like habit of living in trees. In accordance with this change of habit is a relative shortening of the hind-limbs, a feature which begins to be observable in Dorcopsis. "The general build," writes Mr. Thomas, "is of the ordinary mammalian proportions, not macropodiform at all." The muffle is not naked for the greater part, though the shortness of the hairs gives that effect. As in Dorcopsis, but not as in Macropus, the bulla tympani is not swollen. There are altogether five species, the fifth, D. bennetti, having been lately described from specimens living in the Zoological Society's Gardens.