Picea-Pseudotsuga-hylium formation (-ium) Paronychia-Silene-chalicium
Picetum consocies (-etum) Paronychietum
Opulaster-Ribesanum layer (-anum)
Opulasterile society (-ile) Androsacile
Thalictrare community (-are) Festucare
Pirolon family (-on) Arenarion

349. The investigation of a particular formation. A comprehensive and thorough study of a formation should be based upon as many examples of it as are accessible. The example which is at once the most typical and the most accessible is made the base area. This plan saves time and energy, reduces the number of instruments that are absolutely necessary, and establishes a common basis for comparison. The inquiry should be made along four lines, all fundamental to a proper knowledge of the formation. These lines are: (1) the determination of the factors of the habitat, (2) a quadrat and a transect study of the structure of the formation, (3) a similar investigation of development, (4) a floristic study of the contiguous formation, with special reference to migration. The sequence indicated has proven to be the most satisfactory, and is to be regarded as all but absolutely essential. Naturally, this applies only to the order in which the various lines are to be taken up, as they are carried on together when the work is fully under way. Since instrument and quadrat methods have already been given in detail, it is unnecessary that they be repeated. Similarly, the questions which pertain to structure and development and to the surrounding vegetation are considered in detail in the pages which precede.

Fig. 80. Eritrichiare (Eritrichium aretioides), a community of the alpine meadow formation.

CLASSIFICATION AND RELATIONSHIP

350. Bases. Formations may be grouped with reference to habitat or kind, development or position. Classification upon the basis of habitat places together formations which are similar in physiognomy and structure. Developmental classification is based upon the fact that the stages of a particular succession are organically connected or related, though they are normally different in both physiognomy and structure. Grouping with respect to position is made solely upon occurrence in the same division of vegetation. The formations thus brought together usually possess neither similarity of kind or structure, nor do they have any necessary developmental connection. Habitat and developmental classification are of fundamental value; regional arrangement is more superficial in character. All serve, however, to emphasize different relations, and, while the developmental system expresses the most, they should all be used to exhibit the vegetation of a region, province, or zone.

Fig. 81. Pachylophon (Pachylophus caespitosus), a family of the gravel slide formation.

351. Habitat classification. In arranging formations with reference to habitats, the direct factors, water and light, can alone be used to advantage. Such a system is fundamental, because it is founded upon similarity of habitat and of structure. Proposed groupings based upon nutrition-content, or upon the division of factors into climatic and edaphic, have elsewhere[[42]] been shown to be altogether of secondary importance, if not actually erroneous. The basis of the habitat grouping is water-content, which is supplemented by light whenever the factor is decisive. The primary divisions thus obtained are water, forest, grassland, and desert, which are characterized respectively by associations of hydrophytes, mesophytes, hylophytes, poophytes, and xerophytes respectively. Within these, formations are arranged according to the type of habitat, i. e., pond, meadow, forest, dune, etc. These divisions comprise all formations which belong to the type by virtue of their physiognomy and structure. Such formations differ from each other very considerably or completely in the matter of floristic, i. e., component species, but they still belong to the same type. A dune formation in the interior and one on the coast may not have a single species in common, and yet they are essentially alike in habitat, development, and structure.

352. Nomenclature. The names of formations are taken from the habitats which they occupy. Each formation should have a vernacular and a scientific name. The latter is especially important since it ensures brevity and uniformity, and obviates the obscurity and confusion that arise from vernacular terms in many tongues. Scientific names have been made uniformly from Greek words of proper meaning by the addition of the suffix -ium (εῖον), which denotes place.[[43]] The following list gives the English and the scientific name of the various habitats, and their corresponding formations, and indicates the primary divisions into which these fall.