[CHAPTER IV.]
FOSSIL BOTANY.
Fossil Vegetables.—The remains of the vegetable kingdom are presented to the notice of the geologist in various conditions; in some instances these relics are but little changed in their aspect, as, for example, in the recent accumulations of mud and silt, at the bottoms of lakes and rivers, and in morasses, and peat-bogs. In tufaceous incrustations, the imprints of wood, and of leaves and stems, are often sharply defined on the solid masses of concretionary and crystalline limestone.
In the ancient deposits, vegetables are found in two different states. In the one their substance is completely permeated by mineral matter; it may be calcareous (lime), siliceous (flint), ferruginous (iron), or pyritous (sulphuret of iron); and yet both the external characters, and the internal structure, may be preserved. Such are the fossil trees of the Isle of Portland, fragments of which so closely resemble decayed wood, as to deceive the casual observer, until by close examination of their texture and substance he finds that they possess the weight and hardness of stone. In the silicified wood which abounds in many of the tertiary strata, the most delicate tissues of the original are preserved, and by microscopical examination (see [Pl. V.]) may be displayed in a distinct and beautiful manner. In calcareous fossil wood the structure is also retained; and in many limestones, leaves and seed-vessels are well preserved.
The ligneous coverings, or the husks and shells, of nuciferous fruits, and the cones or strobili of Firs and Pines, are frequently met with in an excellent state of preservation; in some rare instances indications of flowers have been observed ([Lign. 67]). The parts of fructification in some of the fern tribe (Lign. [25] and [27]), occur in coal-shale, and in the grit of Tilgate Forest (Wond. p. 394): the pollen, and the resinous secretions of pines and firs, have been discovered in tertiary marls, and in the Greensand. The well-known substance. Amber, so much in request for ornaments, is unquestionably of vegetable origin; it has been found impacted in the trunks of its parent trees (Wond. p. 242). The fossil resin discovered in the London clay, at Highgate and the Isle of Sheppey, is doubtless referable to the coniferæ found in that deposit.
In the Clathrariæ of Tilgate Forest, indications of a resinous secretion have been detected.
The Diamond, which is pure charcoal, is probably a vegetable secretion, that has acquired a crystalline structure by electro-chemical forces. It has been converted into Coke and Graphite by the action of intense heat; and the electrical properties of the substance were changed, the Diamond being an insulator, and the Coke, a conductor of electricity. (Wond. p. 706.)
When the microscope is more extensively employed in investigations of this kind, it is probable that the siliceous spines and stars which begem the foliage of many plants (as the Deutzia, Lithospermum officinale, &c), will be discovered in a fossil state, for they are as indestructible as the frustules of Diatomaceæ, and the spicules of sponges which are so common in flint and chalcedony.
FOSSIL VEGETABLES.
But vegetables occur not only as petrified stems, leaves. and fruits, associated with other remains in the strata, but also in beds of great thickness and extent, consisting wholly of plants transmuted, by that peculiar process which vegetable matter undergoes when excluded from atmospheric influence, and under great pressure, into Lignite, and Coal. And there are intermediate stages of this process, in which the form and structure of the trees and plants are apparent; and a gradual transition may be traced, from the peat-wood and submerged forests of modern epochs, in which leaves, fruits, and trunks of indigenous species are preserved, to those ancient accumulations of carbonaceous matter, whose vegetable origin the eye of science can alone detect.