THE TEST OF INCLUDED ORGANISMS.
The second great law of the Stratigraphical Geologist is that fossiliferous strata are identifiable by their included organisms, in other words, that we can tell the geological age of deposits by examination of the fossils contained in them, though the determination of age must be given in more general terms in some cases than in others. Considerable misconception has arisen concerning the value of fossils as indices of age, and it is necessary therefore to discuss the significance of the law of identification of strata by their included organisms at some length.
The comparison between fossils and medals has frequently been made and fossils have well been styled the "Medals of Creation"; and the significance of fossils as guides to the age of deposits may perhaps be made clearer if we pursue this comparison some way. In the first place there is clear indication of a gradual increase in the complexity of organisation of the fossils as one passes from the earlier to the later rocks, and accordingly the general facies of a fauna is likely to furnish a clue to the age of the rocks in which it is found, even though every species or even genus represented in the fauna was previously unknown to science. So an antiquary versed in the evolution of art or metallurgy, might detect the general age of a medal with whose image and superscription he was not acquainted. He would know that a medal struck in iron was formed subsequently to the bronze age, or that one formed of palladium appertained to the present century. But quite apart from any theoretical knowledge, an antiquary would find as the result of accumulated experience that certain medals are characteristic of certain periods; he would learn that the denarius is characteristic of a different period from that indicated by the coin of the Victorian era, even though he had no knowledge of the technicalities of numismatics. The same is the case with the geologist. He may not be influenced by any knowledge of the evolution of faunas and floras, but actual work amongst the rocks will show him that the trilobite is not found with the belemnite or the ichthyosaur with the elephant, save under exceptional circumstances, which only prove the rule, as for instance when worn bones of ichthyosaurs are washed from their proper strata into gravels with elephant-bones.
It must be distinctly understood that the determination of fossils as characteristic of different periods is solely made as the result of experience. No à priori reasoning may give one indication of the actual range in time of a species or genus; no one can say why Discina has a long range in time, whilst that of the closely related Trematis is very limited. This being the case, the greater the mass of evidence which is accumulated as to the range of a fossil, the greater will be the value of that fossil as a clue to the age of the deposit in which it is found. This is so important, that it requires more than mere notice. If a fossil is found in abundance in a group of strata B in any one area, and is not found in an underlying group A or overlying group C in that area after prolonged search, we may confidently speak of the fossil as characteristic of the strata B in that area, though without further work, the value of the fossil as a clue to age in other areas would be unproved. It may nevertheless happen, that after more prolonged search in A or C, in the original area a few specimens of the fossil which has been spoken of as characteristic of B may be found in one or other of them, in small quantity. The value of the fossil as one characteristic of B will be slightly diminished, though only slightly, as it is not likely to turn up in numbers in the strata A or C after the prolonged search. Should the fossil be found also to be characteristic of the strata B in areas other than the original one, it becomes of more than local value, and if, after much study it is found to characterise the same strata over wide areas, the cumulative evidence now obtained will render the fossil peculiarly important to the stratigraphical geologist. The detection of characteristic fossils is not quite so simple as might be supposed from the above remarks, for examination of the position of one fossil will not prove the contemporaneity of beds in different places, to prove this, all the evidence at our disposal must be considered, for reasons which will be presently pointed out.
As the result of accumulated knowledge, we can now compile lists of characteristic fossils of the major subdivisions of the strata, which are of world-wide utility and as our knowledge increases, we are enabled to subdivide the strata into minor divisions of more than local value.
What is a fossil? Before discussing the value of fossils as aids to the stratigraphical geologist, it may be well to make a few observations as to what constitutes a fossil. It is difficult to give any concise definition, and as is often the case in geology, an explanatory paragraph is of more value than a mere definition. The term fossil was originally applied to anything dug up from the rocks of the earth's crust, and was used with reference to inorganic objects as well as organic remains, for instance minerals were spoken of as fossils. It is now applied essentially though not exclusively to relics of former organisms, though one still reads of fossil rain-drops, fossil sun-cracks, and so on. Furthermore, the relics need not necessarily be parts of the organism, the track of a worm or a bird's nest if embedded in the strata would be termed a fossil. It is generally agreed that no sharp line can be drawn between recent and fossil organic remains which is based upon the degree of mineralisation (or as it was sometimes termed petrifaction) of the relics, for many true fossils have not undergone mineralisation, subsequent to their entombment.
It has been suggested that the name fossil should be applied to organic remains which have been entombed by some process other than human agency, but this restriction is undesirable. The stone-implement of the river gravels is as genuine a fossil as the ammonite extracted from the chalk, and the human relics of very recent date may give information of a character quite similar to that supplied by other remains, for instance, the occurrence of moa-bones in New Zealand in accumulations below those containing biscuit-tins and jam-pots has been used as a geological argument pointing to the extinction of the moa before the arrival of Europeans in New Zealand. The biscuit-tin here serves all the purposes of a fossil, and there is no valid reason why it should not be spoken of as such.
This statement brings one to consider another method which has been adopted in order to separate fossil organisms from recent ones, namely the time-test. This again is inapplicable, for no line can be drawn between the shell which was buried in yesterday's tidal deposit and that which has lain in the strata through geological ages, and each may be equally useful to the geologist.
Whilst, then, we can give no definition of fossil which is likely to meet with general acceptance, the term can be so used, as not to give rise to any doubts as to its meaning, and it is generally applicable to any organic relics which have been embedded in any deposit or accumulation by any agent human or otherwise.
Mode of occurrence of fossils. It will not be out of place to say a few words as to the way in which fossils are found in strata, as beds are often inferred to be unfossiliferous, because of ignorance of methods which should be pursued in searching for organic relics. It is unnecessary to dilate upon the actual modes of preservation of organisms, which is treated of fully in other works. In the first place, it is rash to assert that any deposit is unfossiliferous because no fossils have been found in it, even after prolonged search. The Llanberis slates had been eagerly searched for fossils for many years without result, but that the search was not exhaustive was proved by the discovery of trilobites in them some years ago. Seekers after fossils are rather prone to confine their attention to strata which are already known to be fossiliferous than to pay much attention to those which have hitherto yielded no organic remains.