Some kinds of deposits are more often fossiliferous than others. Limestones which are frequently largely of organic origin, are often rich in remains, and muddy deposits more frequently furnish fossils than those of a purely sandy nature. The difference in the yield is not necessarily due to the original inclusion of more remains in one rock than in another, but is often caused by the obliteration of former relics owing to changes which have taken place in the rocks subsequently to their deposition. No sedimentary rock must be regarded as unfossiliferous, however unfitted it appears for the preservation of fossils. The writer has seen fossils, not only in coarse conglomerates, rocks which frequently contain no traces of organisms, but in deposits composed largely of specular iron ore, and even in intrusive igneous rocks, though in the latter case, the inclusion of fossils was due to circumstances which cannot have occurred with frequency.
In sandy strata, the substance of the fossils has often been completely removed, leaving hollow casts, which may be almost or quite unrecognisable. In these circumstances, much information may be obtained by taking impressions of the casts in modelling wax or some other material. The importance of this process may be judged from the results it yielded to Mr Clement Reid in the case of the fossils of the Pliocene deposits occurring in pipe-like hollows in the Cretaceous rocks of Kent and the discovery of the remarkable reptiles described by Mr E. T. Newton from the Triassic sandstones of Elgin.
In argillaceous rocks which have been affected by the processes producing cleavage, the fossils may be distorted beyond recognition or owing to the difficulty of breaking the rocks along the original planes of deposition, may remain invisible. Under such circumstances, small nodules of sandy or calcareous nature may sometimes be found included in the argillaceous deposits and may perhaps yield fossils. Oftentimes, also, where the argillaceous rock is in close proximity to a harder rock, such as massive grit, the argillaceous rock in close contiguity to the hard rock may escape the impress of cleavage-structure, and fossils may be readily extracted from rocks in this position when not obtainable from other parts of the deposit. It was under these circumstances that the trilobites alluded to above were obtained from the Llanberis slates.
The fossils of calcareous rocks are often very obvious, but difficult to extract, as they break across when the rock is fractured. They are frequently obtainable in a perfect condition when the rock is weathered. Occasionally they may be extracted from certain argillaceous limestones if the limestone be heated to redness, and suddenly plunged into cold water. Fossils are often found in a state which enables them to be readily extracted when a limestone is coarsely crystalline, though they cannot be extracted in a perfect condition when the same limestone is in a different state.
Many microzoa, which are invisible in rocks, even when viewed through a lens, may be found in microscopic sections of calcareous and silicious rocks, and plant structures may be detected under similar circumstances in the case of carbonaceous rocks.
Various special methods of extracting fossils from rocks have been described by different writers, many of which are very complex, and require much time. The mechanical action of the sand-blast and the solvent action of various acids as hydrochloric and hydrofluosilicic have been found of use upon different occasions[11]. The various processes which have been utilised in order to extract and develop fossils can, however, be best learned by information obtainable from curators of palæontological collections, and by actual experience, and there is yet much information to be acquired as to the manner of extracting fossils from various kinds of rocks.
[11] For information concerning use of acids see especially Wiman, C. "Ueber die Graptoliten," Bull. Geol. Inst., Upsala, No. 4, vol. II. Part II.
Relative value of fossils to the Stratigraphical Geologist. It has been hinted above that no general rule as to the relative value of fossils as guides to the age of strata can be laid down, and that the ascertainment of their relative value is largely the result of actual experience. It may be noted, however, that organisms which possess hard parts are naturally more important to the geologist than those which do not, as few traces of the latter are preserved in the fossil state, and even when preserved are usually too obscure to be of much practical use. Of the organisms which do possess hard parts, different groups have been utilised to a different degree, and one group will be more or less important than another, according to the use to which it is applied. Groups of organisms which have a long range in time are naturally useful for the identification of large subdivisions of the strata, whilst those which have had a shorter range are valuable when separating minor subdivisions.
Again, as the bulk of the sedimentary deposits has been formed beneath the waters of the ocean, relics of marine organisms are naturally more useful than those of freshwater ones. Other things being equal, the more easily the organism is recognisable, and the more abundant are its remains, the greater its value to the stratigraphical geologist, and as the remains of invertebrates are usually found in greater quantities and in more readily recognisable condition than those of the vertebrates, they have been used more extensively as indices of age. Of the invertebrates, the mollusca are often very abundant, their remains are adapted for preservation, and their characteristics have been extensively studied, and accordingly they have been and are of great use to the geologist. Of other groups, the graptolites, corals, echinids, brachiopods, and trilobites have been very largely utilised. The Lower Palæozoic strata have been divided into numerous groups, each characterised by definite forms of graptolites, and a similar use has been made of the ammonites in the case of the Mesozoic rocks. It is not to be inferred that these groups of organisms are naturally more useful than other groups, on account of the extent to which they have been used; we can merely state that they have been proved to be useful as the result of prolonged study; when other groups have received equal attention, they may well be found to be equally useful for the purposes which we have in view.
Contemporaneity and Homotaxis. From what has been already stated, it will be recognised that the ages of the various fossiliferous rocks of the geological column[12] in any one area can be identified with greater or less degree of certainty by reference to their included organisms, the various subdivisions being marked by the possession of characteristic fossils, and it will be naturally and rightly inferred that the greater the number of characteristic fossils of any one deposit, the more certain is the identification of that deposit. In practice, geologists are wont to ascertain the age of the strata after consideration of all the fossils found therein, some of which may be actually characteristic whilst many may come up from the strata below, or pass into those above. Having ascertained the order of succession and fossil contents of the strata in various regions, it is the task of the geologist to compare the strata of these two regions, and this task is fraught with considerable difficulty. Much controversy has arisen as to the degree of accuracy with which strata of remote regions can be correlated, and the subject is one which requires full consideration.