When one takes into consideration the fact that, notwithstanding many oscillations of level, the land during Cainozoic times was gradually extending, and the sea disappearing from wide regions which it had formerly covered, one can hardly doubt that the seemingly gradual change from tropical to temperate conditions was due, in large measure, to that persistent continental growth. I confess, however, that it is difficult to account for the very genial climate which continued to prevail over the arctic regions. So far as one can gather from the evidence at present available, some of the marine approaches to those latitudes had been cut off by the movements of elevation which brought the Cainozoic era to a close, while the arctic lands were perhaps more extensive than they are now. The Cretaceous Mediterranean Sea of North America had vanished, and we cannot prove that the Tertiary Sea of southern Europe communicated across the low-grounds of Russia with the Arctic Ocean. We know, however, that the archipelago of southern Europe was in direct connection with the Indian Ocean, and it is most probable that a wide arm of the same sea stretched north from the Aralo-Caspain area through Siberia. Indeed, much of what are now the lowlands of western and northern Asia was probably sea in Tertiary times. It seems likely, therefore, that, even at this late period, marine currents continued to reach the Arctic Zone across the continental plateau. When the warm waters of the Indian Ocean eventually ceased to invade Europe, and the Mediterranean became much restricted in area, the climate of the whole Continent could not fail to be profoundly affected.
There is yet another line of evidence to which brief reference may be made. I have spoken of the remarkable uniformity of climatic conditions which obtained in Palæozoic times, and of the gradual modification of these conditions which subsequently supervened. Now, it is worthy of note that in their lithological characters the oldest sedimentary strata themselves likewise exhibit a prevalent uniformity which in later systems becomes less and less conspicuous. The Cambro-Silurian mechanical sediments, for example, maintain much the same character all the world over; and the like is true, although in a less degree, of the marine accumulations of the Devonian period. The corresponding mechanical deposits of later Palæozoic ages continue to show more and more diversity, but at the same time they preserve a similarity of character over much more extensive areas than is found to be the case with the analogous sediments of the Mesozoic era. Finally, these last are more or less strongly contrasted with the marine mechanical accumulations of Cainozoic times, which are altogether more local in character. This increasing differentiation is quite in keeping with what we know of the evolution of our land-areas. In early Palæozoic ages, when insular conditions prevailed and the major portion of the primeval continental plateau was covered by shallow seas, it is obvious that mechanical sediments would be swept by tidal and other currents over enormous areas, and that these sediments would necessarily assume a more or less uniform character. Indeed, I suspect that much of the sediment of those early seas may have been the result of tidal scour, and that marine erosion was more generally effective then than it is now. With the gradual growth of the land and the consequent deflection and limitation of currents, marine mechanical sediments would tend to become more and more local in character. Thus the increasing differentiation which we observe in passing from the earlier to the later geological systems is just what might have been expected.
Summing up, now, the results of this rapid review of the evidence, we seem justified in coming to the following conclusions:—
(1.) In Palæozoic times, Europe and North America were represented by considerable areas of dry land, massed chiefly in the higher latitudes, while further south groups of smaller islands were scattered over the submerged surface of the primeval continental plateau. The other continents appear, in like manner, to have been represented by islands—some of which may have reached continental dimensions. A very remarkable uniformity of climate accompanied these peculiar geographical conditions.
(2.) In Mesozoic times, the primeval continental plateau came more and more to the surface, but the land-areas were still much interrupted, so that currents from tropical regions continued to have ready access to high latitudes. The climate of the whole globe, therefore, was still uniform, but apparently not so markedly as in the preceding era.
(3.) In Cainozoic times, the land-masses continued to extend, and the sea to retreat from hitherto submerged areas of the continental plateau; and this persistent land-growth was accompanied by a gradual lowering of the temperature of northern and temperate latitudes, and a more and more marked differentiation of climate into zones.
Having thus very briefly sketched the geographical evolution of the land during Palæozoic, Mesozoic, and Tertiary times, and come to the general conclusion that climate has varied according to the relative position of land and sea, I have next to consider the geographical and climatic conditions of the Quaternary period. These, however, are now so well known, that I need to no more than remind you that, so far as the chief features of our lands are considered, all these had come into existence before the dawn of the Ice Age. The greater contours of the surface, which were foreshadowed in Palæozoic times, and which in Mesozoic times were more clearly indicated, had been fully evolved by the close of the Pliocene period. The connection between the Mediterranean and the Indian Ocean probably ceased in late Pliocene times. The most remarkable geographical changes which have taken place since then within European regions have been successive elevations and depressions, in consequence of which the area of our Continent has been alternately increased and diminished. At a time well within the human period, our own islands have been united to themselves and the Continent, and the dry land has extended north-west and north, so as to include Spitzbergen, the Faröe Islands, and perhaps Iceland. On the other hand, our islands have been within a recent period largely submerged. Similarly, in North America, we are furnished with many proofs of like oscillations of level having taken place in Quaternary times. Is it possible, then, to explain the climatic vicissitudes of the Pleistocene period by means of such oscillations? Many geologists have tried to do so, but all these attempts have failed. It is quite true that a general elevation of the land in high latitudes would greatly increase the ice-fields of arctic regions, and might even give rise to perennial snow and glaciers in the mountain-districts of our islands. But it is inconceivable that any such geographical change could have brought about that general lowering of temperature over the whole northern hemisphere which took place in Pleistocene times. For we have to account not only for the excessive glaciation of northern and north-western Europe, and of the northern parts of North America, but for the appearance of snow-fields and glaciers in much more southern latitudes, and in many parts of Asia where no perennial snow now exists. Moreover, we have to remember that arctic conditions of climate obtained in north-western Europe even when the land was relatively much lower than it is at present. The arctic shell-beds of our own and other temperate regions sufficiently prove that geographical conditions were not the only factor concerned in bringing about the peculiar climate of the Pleistocene period. Then, again, we must not forget that at certain stages of the same period genial conditions of climate were coincident with a much wider land-surface in north-western Europe than now exists. The very fact that interglacial deposits occur in every glaciated region is enough of itself to show that the arctic conditions of the Pleistocene could not have resulted entirely from a mere elevation of land in the northern parts of our hemisphere.
The only explanation of the peculiar climatic vicissitudes in question which seems to meet the facts, so far as these have been ascertained, is the well-known theory advanced by Dr. Croll. After carefully considering all the objections which have been urged against that theory, there is only one, as it seems to me, that is deserving of serious attention. This objection is not based on any facts connected with the Pleistocene deposits themselves, but on evidence of quite another kind. It is admitted that were the Pleistocene deposits alone considered, Croll’s theory would fully account for the phenomena. But, it is argued, we cannot take the Pleistocene by itself, for if that theory be true, then climatic conditions similar to those of the Pleistocene must have supervened again and again during the past. Where, then, we are asked, is there any evidence in Palæozoic, Mesozoic, or Cainozoic strata of former widespread glacial conditions? If continental ice-sheets, comparable to those of the Pleistocene, ever existed in the earlier ages, surely we ought to find more or less unmistakable traces of them. Now, at first sight, this looks a very plausible objection, but it has always seemed to me to be based upon an assumption that is not warranted by our knowledge of geographical evolution. Dr. Croll always admitted implicitly that high eccentricity of the earth’s orbit might have happened again and again without inducing glacial conditions like those of the Pleistocene. The objection takes no account of the fact that the excessive climate of the Glacial period was only possible because of special geographical conditions—conditions that do not appear to have been fully evolved before Pliocene times. No one has seen this more clearly than Mr. Wallace,[DL] with the general drift of whose argument I am quite at one. In earlier ages, the warm water of the tropics overflowed wide areas of our present continents—most of the dry land was more or less insular, and the seas within the Arctic Circle were certainly not cold as at present, but temperate and even genial. If we go back to Cambro-Silurian times, we find only the nuclei, as it were, of our existing continents appearing above the surface of widespread shallow seas. It is quite impossible, therefore, that under such geographical conditions, great continuous ice-sheets, like those of the Pleistocene, could have existed—no matter how high the eccentricity of the earth’s orbit may have been. The most that could have happened during such a period of eccentricity would be the accumulation of snow-fields on mountains and plateaux of sufficient height, the formation here and there of local glaciers, and the descent of these in some places to the sea. And what evidence of such local glaciation might we now expect to find? No old land-surface of that far-distant period has come down to us: we look in vain for Cambro-Silurian roches moutonnées and boulder-clay or moraines. The only evidence we could expect is just that which actually occurs, namely, erratics (some of them measuring five feet and more in diameter) embedded in marine deposits. It may be said that a few erratics are hardly sufficient to prove that a true Glacial period supervened in Cambro-Silurian times, and I do not insist that they are. But I certainly maintain that if any lowering of the temperature were induced by high eccentricity of the earth’s orbit during Cambro-Silurian times, then ice-floated erratics are the only evidence of refrigeration that we need ever hope to find. The geographical conditions of early Palæozoic times forbade the formation of enormous ice-sheets like those of the Pleistocene period. Extreme climatic changes were then impossible, and periods of high eccentricity might have come and gone without inducing any modifications of flora and fauna which we could now recognise. We are ignorant of the terrestrial life of the globe at that distant period, and our knowledge of the marine fauna is not sufficient to enable us to deny the possibility of moderate fluctuations in the temperature of the seas of early Palæozoic times. Moreover, we must not forget there were then no such barriers to migration as now exist. If the conditions became temporarily unsuitable, marine organisms were free to migrate into more genial waters, and to return to their former habitats when the unfavourable conditions had passed away.
[DL] See Island Life.
The uniform climate so characteristic of the Cambro-Silurian period appears to have prevailed likewise during the later stages of the Palæozoic era. This we gather from a general consideration of the floras and faunas, and their geographical distribution. The dry land, as we have seen, continued to increase in extent; but vast areas of the primeval continental plateau of the globe still continued under water, and currents from southern latitudes flowed unrestricted into polar regions. During the protracted lapse of time required for the formation of the later Palæozoic systems several periods of high eccentricity must have occurred. But, so far as one can judge, the disposition of the larger land-areas was never such as to induce a true Ice Age. Nevertheless we are not without evidence of ice-action in Old Red Sandstone, Carboniferous, and Permian strata. And it seems to me probable that the erratic accumulations referred to may really indicate local action, of more or less intensity, brought about by such lowering of the temperature as would supervene during a period of high eccentricity. It is true we may explain the phenomena by inferring the existence of mountains of sufficient elevation—and this, indeed, is the usual explanation. But it is doubtful whether those who adopt that view have fully considered what it involves. Take, for example, the case of the breccias and conglomerates of the Lammermuir Hills, which have all the appearance of being glacial and fluvio-glacial detritus. These deposits overlie the highly-denuded Silurian greywackés of Haddingtonshire in the north and of Berwickshire in the south, and have evidently been derived from the intervening high-grounds—the width of which between the Old Red Sandstone accumulations in question does not exceed eight or nine miles. The breccias reach a height of 1300 feet, while the dominating point of the intervening uplands is 1700 feet. Under present geographical conditions it is doubtful whether perennial snow and glaciers of any size at all could exist in the region of the Lammermuirs at a less altitude than 7000 feet or more. But between the breccias of Haddingtonshire and the equivalent deposits in Berwickshire there is no space for any intermediate range of mountains of circumdenudation of such a height. Moreover, we must remember that under the extremely uniform conditions which obtained in Palæozoic times the snow-line could not possibly have been attained even at that elevation. When the Devonian coral-reefs described by Dupont were growing in the sea that overflowed western Europe, to what height must the southern uplands of Scotland have been elevated in order to reach the snow-line! We may make what allowance we choose for the denudation which the Silurian rocks of the Lammermuirs must have experienced since the deposition of the Old Red Sandstone, but it is simply a physical impossibility that mountains of circumdenudation of the desiderated height could ever have existed in the Lammermuir region at the time the coarse breccias were being accumulated.[DM] It seems to me, then, that these breccias are in every way better accounted for by a lowering of temperature due to increased eccentricity of the orbit. This view frees us from the necessity of postulating excessive upheavals over very restricted areas, and of creating Alps where no Alps could have existed.