But, though it may at first sight have appeared that an admission of the doctrine of catastrophes is in harmony with a providential government of the world, and that the emergence of different organic forms in successive ages is a manifestation of creative intervention, of which it was admitted that as many as from twelve to twenty, if no more, successive instances might be recognized, we may well congratulate ourselves that those important doctrines rest upon a far more substantial basis. Rightly considered, the facts lead to a very different conclusion. Successive forms assumed by man. Physiological investigations have proved that all animals, even man, during the process of development, pass in succession through a definite cycle of forms. Starting from a simple cell, form after form, in a definite order is assumed. In this long line of advance the steps are ever, in all individuals, the same. But no one would surely suppose that the changed aspect at any moment presented is due to a providential interposition. But they are rigidly determined by law. On the contrary, it is the inevitable result of what has been taking place under the law of development, and the sure precursor of what is about to follow. In the organic world, the successive orders, and genera, and species are the counterparts of these temporary embryonic forms of the individual. Indeed, we may say of those successive geological beings that they are mere embryos of the latest—embryos that had gained a power of reproduction. How shall we separate the history of the individual from the history of the whole? Do not the fortunes and way of progress of the one follow the fortunes and way of progress of the other? If, in a transitory manner, these forms are assumed by the individual, equally in a transitory manner are they assumed by the race. Nor would it be philosophical to suppose that the management in the one instance differs from the management in the other. If the one is demonstrably the issue of a law in action, so [325] must the other be too. It does not matter that the entire cycle is passed through by the individual in the course of a few months, while in the race it demands ages. Individual and race development conducted in the same way. The standard of time that ought to be applied is the respective duration of life. In man it is much if he attains to threescore years and ten; but the entire period of human record, embracing several thousand years, offers not a single instance of the birth, maturity, and death of a species. They, therefore, who think they find, in the successive species that have in an orderly manner replaced each other in the life of the earth, the sure proof of Divine intervention, would do well to determine at what point the production of such forms by law ceases, and at what point their production by the immediate act of God begins. Their task will be as hard to tell where one colour in the rainbow ends and where the next commences. They will also do well to remember that, in great mundane events, the scale of time is ample, and that there may be no essential difference between a course that is run over in a few days and one that requires for its completion thousands of centuries.

Catastrophes disproved by the co-existence of types.The co-existence of different types in the organic series was the incontrovertible fact by which was demonstrated the gradual passage from form to form without catastrophes, the argument relied upon gathering strength from such circumstances as these, that even the fossil shells of the modern Italian tuffs which are not extinct exhibit a slight want of correspondence when compared with those now inhabiting the Mediterranean, some of the old ones being twice and a half as large as the present, and that there is a numerical passage from strata containing seventy per cent. of recent shells to those that are altogether recent, or contain one hundred per cent. This is manifestly indicative of a continually changing impression bringing on a corresponding modelling. It is the proof of a slow merging into, or of a measured assumption of, the new form—a transition, for the completion of which probably a very long time is required. That the existing reindeer is found in the same fluviatile deposits with an extinct hippopotamus seemed certainly to prove that there was a condition of things [326] in which the co-life of those animals was possible in the same locality, and that, as the physical causes slowly changed, the one might be eliminated and the other might be left. That the regulating conditions were altogether physical was obvious from such facts as that in the bone-caves of Australia all the mammals are marsupial, and in the pampas of South America they are allied to such forms as are indigenous, armadilloes, sloths, etc., showing the tokens of lineage or hereditary transmission. For still more remote times numerous instances of a similar nature were detected; thus, throughout the whole Secondary period, the essential characteristic was the wonderful development of reptile life, while in the Tertiary it was the development of mammals. But the appearance of mammals had commenced long before that of reptiles had ceased. Indeed, the latter event is incomplete in our times; for, though the marine Saurians have been almost entirely removed, the fluviatile and terrestrial ones maintain themselves, though diminished both in species and individuals. Now such an overlapping of reptiles and mammals was altogether irreconcilable with the doctrine of a crisis or catastrophe, and, in fact, it demonstrated the changing of organisms in the changing of physical states.

Cuvier's doctrine of permanence of species.Cuvier maintained the doctrine of the permanence of animal species from the facts that the oldest known do not appear to have undergone any modification, and that every existing one shows a resistance to change. If his observations are restricted to periods not exceeding human history, they may perhaps be maintained, but that duration cannot be looked upon as more than a moment in the limitless progress we are considering, and it was in this view that Cuvier's doctrine proved to be incapable of defence. Imperfection of evidence in its support. What does it signify if our domestic animals show no variations when compared with the corresponding images depicted on the hieroglyphic monuments of Egypt, or with the descriptions left by ancient authors? Evidence of that kind is valueless. Does the geologist ask of the architect his opinion whether there have ever been upliftings and down-sinkings of the earth? If he did, would not every structure in Europe be [327] brought forward as an evidence that nothing of the kind had ever occurred? A leaning tower, or a church with inclining walls in Italy, might pass for nothing; the Pyramids would testify that Egypt itself had never undergone any disturbance—they remain solid on their bases, undisturbed. But what is the weight of all this when placed in opposition with the mass of evidence offered by inclined and fractured strata? And yet such is precisely the proof offered in behalf of the permanence of animals. The facts with which the zoologist deals, like those on which the architect depends, are insufficient for the purpose—they are wanting in extent of time. There have been movements in the crust of the earth, though every building in the world may be perpendicular; there have been transformations of organisms, though for four thousand years there may have been no perceptible change.

Control of organisms by physical conditions.If ever there had been a universal creation of all possible organic forms or combinations, forthwith vast numbers of them must have disappeared, every type being eliminated which was not in correspondence with the external conditions or with the medium in which it was placed. If the environment or the physical conditions underwent a variation, a corresponding variation in the forms that could by possibility exist must ensue, and, from a thorough study of those not eliminated, the physical conditions might be ascertained; and conversely, from a thorough knowledge of the physical conditions, the forms that could escape elimination might be designated. The facts on which Cuvier rested did not demonstrate what he supposed. His immobility of species was no consequence of an innate or intrinsic resistance possessed by them, but merely an illustration that external physical agents had not undergone any well-marked variation in the time with which he was concerned.

Nature of variation of physical conditions.What is here meant by variation in physical forces or condition is not any intrinsic change in their nature, but the varied manner in which they may work by interfering with one another, or experiencing declines of intensity. From the fact that we may read in the fixed stars, through the progressive motion of light, the history of a million of [328] past years, we may be sure that the forces of nature have undergone no intrinsic change; that light was propagated at the same rate, was capable of producing the same optical and chemical effects, and varied in its intensity by distance as it does now; that heat determined corporeal magnitudes. These are things that in their nature are absolutely unchangeable. Always, as now, the freezing of water, and its boiling under a given pressure, must have been the same; there must have been a thermometric zero of life and an upward limit, no animal process ever going on below 32° Fahrenheit or above 212° Fahrenheit.

Effect thereof on organisms.But out of this invariability of natural causes variations in their condition of action arise, and it is these that affect organic forms. Of such forms, some become at length incapable of maintaining themselves in the slow progress of change; others acclimatize, or accommodate, or suit themselves thereto by undergoing modifications, and this was at last discerned to be the true explanation of extinctions and appearances, events taking place very slowly in untold periods of time, and rather by imperceptible degrees than by a sudden catastrophe or crisis.

Transmutation of species.The doctrine of the transmutation of species has met with no little resistance. They who have refused to receive it as one of the truths of Nature have perhaps not given full weight to physiological evidence. When they ask, Has any one ever witnessed such an event as the transmutation of one species into another? has any experimenter ever accomplished it by artificial means? they do not take a due account of time. In the Fables it is related that when the flowers were one evening conversing, "Our gardener," said the rose to the lily, "will live for ever. I have not seen any change in him. The tulip, who died yesterday, told me that she had remarked the same thing; she believed that he must be immortal. I am sure that he never was born."

Two modes of action.Two modes have been presented by which we may conceive of the influence of physical agents upon organic forms. Their long persistent action upon the individual may give rise to modifications, developing one part, stunting another; and such variations, being transmitted in an hereditary way, may become [329] firmly fixed at last. Thus a given plant may, in the course of ages, under the influence of unremittingly acting physical conditions, undergo a permanent change, and a really new plant arise as soon as, through the repetitions of successive generations, the modifications have become so thorough, so profound, as to be capable of transmission with certainty. Perhaps this is what has taken place with many of our kitchen-garden plants, of which the special varieties may be propagated by seeds. But there is another mode by which that result may be reached, even if we decline the doctrine of St. Augustine, who, in his work "De Civitate Dei," shows how islands may be peopled with animals by "spontaneous generation." All organic forms originally spring from a simple cell, the development of which, as indicated by the final form attained, is manifestly dependent on the physical conditions it has been exposed to during its course. If those conditions change, that final form must change correspondingly; and in this manner, since all organic beings come from the same starting-point—the same cell, as has been said, which helplessly submits to whatever impression may be put upon it—the issue is the same as though a transformation or transmutation had occurred, since the descendant is not like its ancestors. Such a manner of considering these changes is in harmony with our best physiological knowledge, since it does not limit itself to a small portion of the life of an individual, but embraces its whole cycle or career. For the more complete examination of this view I may refer to the second chapter of the second book of my "Physiology."

Problem of the modification of forms.But here has arisen the inquiry, Does the modification of organic forms depend exclusively on the impressions of external influences, or is it due to a nisus or force of development residing in the forms themselves?

Whether we consider the entire organic series in its succession, or the progress of an individual in his development, the orderly course presented might seem to indicate that the operation is taking place under a law—an orderly progression being always suggestive of the operation of law. But a philosophical caution must, however, be here exercised; for deceptive appearances [330] may lead us into the error of imputing to such a law, impressed by the Creator on the developing organism, that which really belongs to external physical conditions, which, on their part, are following a law of their own. What is here meant may be illustrated by the facts that occur on the habitable surface of a planet suffering a gradual decline of heat. Three solutions of it. On such a surface a succession of vegetable types might make its appearance, and, as these different types emerged or were eliminated, we might speak of the events as creations and extinctions, and therefore as the acts of God. Or, in the second place, we might refer them to an intrinsic force of development imparted to each germ, which reached in due season its maximum, and then declined and died out; and, comparing each type with its preceding and succeeding ones, the interrelation might be suggested to us of the operation of a controlling law. Or, in the third place, we might look to the external physical condition—the decline of heat—itself taking place at a determinate rate under a mathematical law, and drawing in its consequences the organic variations observed.