Fig. 121.—Perfect Ammonite.
XIX.—Fig. 1.—Apiocrinites rotundus. Fig. 2.—Encrinus liliiformis.
Among the Echinoderms, Cidaris glandiferus, Apiocrinus Roissyanus, and A. rotundus, the graceful Saccocoma pectinata, Millericrinus nodotianus, Comatula costata, and Hemicidaris crenularis may be mentioned; Apiocrinites rotundus, figured in [Plate XIX.], is a reduced restoration: 1, being expanded; a, closed; 3, a cross section of the upper extremity of the pear-shaped head; 4, a vertical section showing the enlargement of the alimentary canal, with the hollow lenticular spaces which descend through the axis of the column, forming the joints, and giving elasticity and flexure to the whole stem, without risk of dislocation. A. rotundus is found at Bradford in Wiltshire, Abbotsbury in Dorset, at Soissons, and Rochelle. This species—known as the Bradford Pear-Encrinite—is only found in the strata mentioned.
The Corals of this epoch occur in great abundance. We have already remarked that these aggregations of Polyps are often met with at a great depth in the strata. These small calcareous structures have been formed in the ancient seas, and the same phenomenon is extending the terrestrial surface in our days in the seas of Oceania, where reefs and atolls of coral are rising by slow and imperceptible steps, but with no less certainty. Although their mode of production must always remain to some extent a mystery, the investigations of M. Lamaroux, Mr. Charles Darwin, and M. D’Orbigny have gone a long way towards explaining their operations; for the Zoophyte in action is an aggregation of these minute Polyps. Describing what he believes to be a sea-pen, a Zoophyte allied to Virgularia Patagonia, Mr. Darwin says: “It consists of a thin, straight, fleshy stem, with alternate rows of polypi on each side, and surrounding an elastic stony axis. The stem at one extremity is truncate, but at the other is terminated by a vermiform fleshy appendage. The stony axis which gives strength to the stem, may be traced at this extremity into a mere vessel filled with granular matter. At low water hundreds of these zoophytes might be seen, projecting like stubble, with the truncate end upwards, a few inches above the surface of the muddy sand. When touched or pulled, they drew themselves in suddenly, with force, so as nearly or quite to disappear. By this action, the highly-elastic axis must be bent at the lower extremity, where it is naturally slightly curved; and I imagine it is by this elasticity alone that the zoophyte is enabled to rise again through the mud. Each polypus, though closely united to its brethren, has a distinct mouth, body, and tentacula. Of these polypi, in a large specimen there must be many thousands. Yet we see that they act by one movement; that they have one central axis, connected with a system of obscure circulation.” Such is the brief account given by a very acute observer of these singular beings. They secrete the calcareous matter held in solution in the oceanic waters, and produce the wonderful structures we have now under consideration; and these calcareous banks have been in course of formation during many geological ages. They just reach the level of the waters, for the polyps perish as soon as they are so far above the surface that neither the waves nor the flow of the tides can reach them. In the Oolitic rocks these banks are frequently found from twelve to fifteen feet thick, and many leagues in length, and preserving, for the most part, the relative positions which they occupied in the sea while in course of formation.
The rocks which now represent the Middle Oolitic Period are usually divided into the Oxford Clay, the lower member of which is an arenaceous limestone, known as the Kellaways Rock, which in Wiltshire and other parts of the south-west of England attains a thickness of eight or ten feet, with the impressions of numerous Ammonites, and other shells. In Yorkshire, around Scarborough, it reaches the thickness of thirty feet; and forms well-developed beds of bluish-black marl in the department of Calvados, in France. It is the base of this clay which forms the soil (Argile de Dives) of the valley of the Auge, renowned for its rich pasturages and magnificent cattle. The same beds form the base of the oddly-shaped but fine rocks of La Manche, which are popularly known as the Vaches Noires (or black cows)—a locality celebrated, also, for its fine Ammonites transformed into pyrites.
The Oxford Clay constitutes the base of the hills in the neighbourhood of Oxford, forming a bed of clay sometimes more than 600 feet thick. It is found well-developed in France, at Trouville, in the department of the Calvados; and at Neuvisy, in the department of the Ardennes, where it attains a thickness of about 300 feet. It is a bluish, sometimes whitish limestone (often argillaceous), and bluish marl. The Gryphæa dilatata is the most common fossil in the Oxford Clay. The Coral Rag is so called from the fact that the limestone of which it is chiefly composed consists, in part, of an aggregation of considerable masses of petrified Corals; not unlike those now existing in the Pacific Ocean, supposing them to be covered up for ages and fossilised. This coral stratum extends through the hills of Berkshire and North Wilts, and it occurs again near Scarborough. In the counties of Dorset, Bedford, Buckingham, and Cambridge, and some other parts of England, the limestone of the Coral Rag disappears and is replaced by clay—in which case the Oxford Clay is overlaid directly by the Kimeridge Clay. In France it is found in the departments of the Meuse, of the Yonne, of the Ain, of the Charente Inférieure. In the Alps the Diceras limestone is regarded, by most geologists, as coeval with the English Coral Rag.