Fig. 145.
a, skull of Monitor Niloticus; b, under-jaw of same.
The masterly memoir of Cuvier, while confirming all the views of Camper, has restored the individuality of this surprising being, which has since received the name of Mosasaurus, that is to say, Saurian or Lizard of the Meuse. It appears, from the researches of Camper and Cuvier, that this reptile of the ancient world formed an intermediate genus between the group of the Lacertilia, which comprehends the Monitors (represented in [Fig. 145]), and the ordinary Lizards; and the Lacertilia, whose palates are armed with teeth, a group which embraces the Iguana and the Anolis. In respect to the Crocodiles, the Mosasaurus resembles them in so far as they all belong to the same class of Reptiles.
The idea of a lizard, adapted for living and moving with rapidity at the bottom of the water, is not readily conceived; but a careful study of the skeleton of the Mosasaurus reveals to us the secret of this anatomical mechanism. The vertebræ of the animal are concave in front and convex behind; they are attached by means of orbicular or arched articulations, which permitted it to execute easily movements of flexion in any direction. From the middle of the back to the extremity of the tail these vertebræ are deficient in the articular processes which support and strengthen the trunk of terrestrial vertebrated animals: they resemble in this respect the vertebræ of the Dolphins; an organisation necessary to render swimming easy. The tail, compressed laterally at the same time that it was thick in a vertical direction, constituted a straight rudder, short, solid, and of great power. An arched bone was firmly attached to the body of each caudal vertebra in the same manner as in Fishes, for the purpose of giving increased power to the tail; finally, the extremities of the animal could scarcely be called feet, but rather paddles, like those of the Ichthyosaurus, the Plesiosaurus, and the Whale. We see in [Fig. 146] that the jaws are armed with numerous teeth, fixed in their sockets by an osseous base, both large and solid. Moreover, an altogether peculiar dental system occupies the vault of the palate, as in the case of certain Serpents and Fishes, where the teeth are directed backwards, like the barb of a hook, thus opposing themselves to the escape of prey. Such a disposition of the teeth sufficiently proves the destructive character of this Saurian.
Fig. 146.—Head of Mosasaurus Camperi.
The dimensions of this aquatic lizard, estimated at twenty-four feet, are calculated to excite surprise. But, as we have already seen, the Ichthyosauri and Teleosauri were of great dimensions, as were also the Iguanodon and Megalosaurus, which were ten times the size of living Iguanas. In all these colossal forms we can only see a difference of dimensions, the aggrandisement of a type; the laws which affected the organisation of all these beings remain unchanged, they were not errors of Nature—monstrosities, as we are sometimes tempted to call them—but simply types, uniform in their structure, and adapted by their dimensions to the physical conditions with which God had surrounded them.