Fig. 184.—Head of Hyæna spelæa.
The Hyænas of our age consist of two species, the striped and the spotted Hyænas. The last presents considerable conformity in its structure with that of the Post-pliocene period, which Cuvier designates under the name of the fossil Spotted Hyæna. It seems to have been only a little larger than the existing species. [Fig. 184] represents the head of the Hyæna spelæa, whose remains, with those of others, were found in the caves of Kirkdale and Kent’s Hole; the remains of about 300 being found in the former. Dr. Buckland satisfied himself, from the quantity of their dung, that the Hyænas had lived there. In the cave were found remains of the ox, young elephant, rhinoceros, horse, bear, wolf, hare, water-rat, and several birds. All the bones present an appearance of having been broken and gnawed by the teeth of the Hyænas, and they occur confusedly mixed in loam or mud, or dispersed through the crust of stalagmite which covered the contents of the cave.
The Horse dates from the Quaternary epoch, if not from the last period of the Tertiary epoch. Its remains are found in the same rocks with those of the Mammoth and the Rhinoceros. It is distinguished from our existing Horse only by its size, which was smaller—its remains abound in the Post-pliocene rocks, not only in Europe, but in America; so that an aboriginal Horse existed in the New World long before it was carried thither by the Spaniards, although we know that it was unknown at the date of their arrival. “Certainly it is a marvellous fact in the history of the Mammalia, that in South America, a native horse should have lived and disappeared, to be succeeded in after ages by the countless herds descended from the few introduced with the Spanish colonists!”[99]
The Oxen of the period, if not identical with, were at least very near to our living species. There were three species: the Bison priscus, B. primigenius, and B. Pallasii; the first with slender legs, with convex frontal, broader than it was high, and differing but slightly from the Aurochs, except in being taller and by having larger horns. The remains of Bison priscus are found in England, France, Italy, Germany, Russia, and America. Bison primigenius was, according to Cuvier, the source of our domestic cattle. The Bos Pallasii is found in America and in Siberia, and resembles in many respects the Musk-ox of Canada.
Where these great Mammals are found we generally discover the fossil remains of several species of Deer. The palæontological question as regards these animals is very obscure, and it is often difficult to determine whether the remains belong to an extinct or an existing species. This doubt does not extend, however, to the gigantic forest-stag, Cervus megaceros, one of the most magnificent of the antediluvian animals, whose remains are still frequently found in Ireland in the neighbourhood of Dublin; more rarely in France, Germany, Poland, and Italy. Intermediate between the Fallow-deer and the Elk, the Cervus megaceros partakes of the Elk in its general proportions and in the form of its cranium, but it approaches the Fallow-deer in its size and in the disposition of its horns. These magnificent appendages, however, while they decorated the head of the animal and gave a most imposing appearance to it, must have sadly impeded its progress through the thick and tangled forests of the ancient world. The length of these horns was between nine and ten feet; and they were so divergent that, measured from one extremity to the other, they occupied a space of between three and four yards.
The skeleton of the Cervus megaceros is found in the deposits of calcareous tufa, which underlie the immense peat moss of Ireland; sometimes in the turf itself, as near the Curragh in Kildare; in which position they sometimes occur in little mounds piled up in a small space, and nearly always in the same attitude, the head aloft, the neck stretched out, the horns reversed and thrown downwards towards the back, as if the animal, suddenly immersed into marshy ground, had been under the necessity of throwing up its head in search of respirable air. In the Geological Cabinet of the Sorbonne, at Paris, there is a magnificent skeleton of Cervus megaceros; another belongs to the College of Surgeons in London; and there is a third at Vienna.
The most remarkable creatures of the period, however, were the great Edentates—the Glyptodon, the gigantic Megatherium, the Mylodon and the Megalonyx. The order of Edentates is more particularly characterised by the absence of teeth in the fore part of the mouth. The masticating apparatus of the Edentates consists only of molars, the incisors and canine teeth being, with a few exceptions, absent altogether, as the animals composing this order feed chiefly on insects or the tender leaves of plants. The Armadillo, Anteater and Pangolin, are the living examples of the order. We may add, as still further characteristics, largely developed claws at the extremities of the toes. The order seems thus to establish itself as a zoological link in the chain between the hoofed Mammals and the ungulated animals, or those armed with claws. All these animals are peculiar to the continent of America.
The Glyptodon, which appears during the Quaternary period, belonged to the family of Armadilloes, and their most remarkable feature was the presence of a hard, scaly shell, or coat of mail six feet in length, and composed of numerous segments, which covered the entire upper service of the animal from the head to the tail. It was, in short, a mammiferous animal, which appears to have been enclosed in a shell like that of a Turtle; it resembled in many respects the Dasypus or Anteater, and had sixteen teeth in each jaw. These teeth were channelled laterally with two broad and deep grooves, which divided the surface of the molars into three parts, whence it was named the Glyptodon. The hind feet were broad and massive, and evidently designed to support a vast incumbent mass; it presented phalanges armed with short thick and depressed nails or claws. The animal was, as we have said, enveloped in, and protected by, a cuirass, or solid carapace, composed of plates which, seen from beneath, appeared to be hexagonal and united by denticulated sutures: above they represented double rosettes. The habitat of Glyptodon clavipes was the pampas of Buenos Ayres, and the banks of an affluent of the Rio Santo, near Monte Video; specimens have been found not less than nine feet in length.
The tesselated carapace of the Glyptodon was long thought to belong to the Megatherium; but Professor Owen shows, from the anatomical structure of the two animals, that the cuirass belonged to one of them only, namely, the Glyptodon.