Another hypothesis, which has been advanced with much less certainty than the preceding, relates to the chemical composition of the air during the Carboniferous period. Seeing the enormous mass of vegetation which then covered the globe, and extended from one pole to the other; considering, also, the great proportion of carbon and hydrogen which exists in the bituminous matter of coal, it has been thought, and not without reason, that the atmosphere of the period might be richer in carbonic acid than the atmosphere of the present day. It has even been thought that the small number of (especially air-breathing) animals, which then lived, might be accounted for by the presence of a greater proportion of carbonic acid gas in the atmosphere than is the case in our own times. This, however, is pure assumption, totally deficient in proof. Nothing proves that the atmosphere of the period in question was richer in carbonic acid than is the case now. Since we are only able, then, to offer vague conjectures on this subject, we cannot profess with any confidence to entertain the opinion that the atmospheric air of the Carboniferous period contained more carbonic acid gas than that which we now breathe. What we can remark, with certainty, as a striking characteristic of the vegetation of the globe during this phase of its history, was the prodigious development which it assumed. The Ferns, which in our days and in our climate, are most commonly only small perennial plants, in the Carboniferous age sometimes presented themselves under lofty and even magnificent forms.
Fig. 38.—Calamite restored. Thirty to forty feet high.
Every one knows those marsh-plants with hollow, channelled, and articulated cylindrical stems; whose joints are furnished with a membranous, denticulated sheath, and which bear the vulgar name of “mare’s-tail;” their fructification forming a sort of catkin composed of many rings of scales, carrying on their lower surface sacs full of spores or seeds. These humble Equiseta were represented during the Coal-period by herbaceous trees from twenty to thirty feet high and four to six inches in diameter. Their trunks, channelled longitudinally, and divided transversely by lines of articulation, have been preserved to us: they bear the name of Calamites. The engraving ([Fig. 38]) represents one of these gigantic mare’s-tails, or Calamites, of the Coal-period, restored under the directions of M. Eugene Deslongchamps. It is represented with its fronds of leaves, and its organs of fructification. They seem to have grown by means of an underground stem, while new buds issued from the ground at intervals, as represented in the engraving.
The Lycopods of our age are humble plants, scarcely a yard in height, and most commonly creepers; but the Lycopodiaceæ of the ancient world were trees of eighty or ninety feet in height. It was the Lepidodendrons which filled the forests. Their leaves were sometimes twenty inches long, and their trunks a yard in diameter. Such are the dimensions of some specimens of Lepidodendron carinatum which have been found. Another Lycopod of this period, the Lomatophloyos crassicaule, attained dimensions still more colossal. The Sigillarias sometimes exceeded 100 feet in height. Herbaceous Ferns were also exceedingly abundant, and grew beneath the shade of these gigantic trees. It was the combination of these lofty trees with such shrubs (if we may so call them), which formed the forests of the Carboniferous period. The trunks of two of the gigantic trees, which flourished in the forests of the Carboniferous period, are represented in [Figs. 39] and [40], reduced respectively to one-fifth and one-tenth the natural size.
What could be more surprising than the aspect of this exuberant vegetation!—these immense Sigillarias, which reigned over the forest! these Lepidodendrons, with flexible and slender stems! these Lomatophloyos, which present themselves as herbaceous trees of gigantic height, furnished with verdant leaflets! these Calamites, forty feet high! these elegant arborescent Ferns, with airy foliage, as finely cut as the most delicate lace! Nothing at the present day can convey to us an idea of the prodigious and immense extent of never-changing verdure which clothed the earth, from pole to pole, under the high temperature which everywhere prevailed over the whole terrestrial globe. In the depths of these inextricable forests parasitic plants were suspended from the trunks of the great trees, in tufts or garlands, like the wild vines of our tropical forests. They were nearly all pretty, fern-like plants—Sphenopteris, Hymenophyllites, &c.; they attached themselves to the stems of the great trees, like the orchids and Bromeliaceæ of our times.
Fig. 39.—Trunk of Calamites. One-fifth natural size.