The Permian rocks have of late years assumed great interest, particularly in England, in consequence of the evidence their correct determination affords with regard to the probable extent, beneath them, of the coal-bearing strata which they overlie and conceal; thus tending to throw a light upon the duration of our coal-fields, one of the most important questions of the day in connection with our industrial resources and national prosperity.
On the opposite page an ideal view of the earth during the Permian period is represented ([Pl. XII.]). In the background, on the right, is seen a series of syenitic and porphyritic domes, recently thrown up; while a mass of steam and vapour rises in columns from the midst of the sea, resulting from the heat given out by the porphyries and syenites. Having attained a certain height in the cooler atmosphere, the columns of steam become condensed and fall in torrents of rain. The evaporation of water in such vast masses being necessarily accompanied by an enormous disengagement of electricity, this imposing scene of the primitive world is illuminated by brilliant flashes of lightning, accompanied by reverberating peals of thunder. In the foreground, on the right, rise groups of Tree-ferns, Lepidodendra, and Walchias, of the preceding period. On the sea-shore, and left exposed by the retiring tide, are Molluscs and Zoophytes peculiar to the period, such as Producta, Spirifera, and Encrinites; pretty plants—the Asterophyllites—which we have noticed in our description of the Carboniferous age, are growing at the water’s edge, not far from the shore.
During the Permian period the species of plants and animals were nearly the same as those already described as belonging to the Carboniferous period. Footprints of reptilian animals have been found in the Permian beds near Kenilworth, in the red sandstones of that age in the Vale of Eden, and in the sandstones of Corncockle Moor, and other parts of Dumfriesshire. These footprints, together with the occurrence of current-markings or ripplings, sun-cracks, and the pittings of rain-drops impressed on the surfaces of the beds, indicate that they were made upon damp surfaces, which afterwards became dried by the sun before the flooded waters covered them with fresh deposits of sediment, in the way that now happens during variations of the seasons in many salt lakes.[51] M. Ad. Brongniart has described the forms of the Permian flora as being intermediate between those of the Carboniferous period and of that which succeeds it.
Although the Permian flora indicates a climate similar to that which prevailed during the Carboniferous period, it has been pointed out by Professor Ramsay, as long ago as 1855, that the Permian breccia of Shropshire, Worcestershire, &c., affords strong proofs of being the result of direct glacial action, and of the consequent existence at the period of glaciers and icebergs.
That such a state of things is not inconsistent with the prevalence of a moist, equable, and temperate climate, necessary for the preservation of a luxuriant flora like that of the period in question, is shown in New Zealand; where, with a climate and vegetation approximating to those of the Carboniferous period, there are also glaciers at the present day in the southern island.
Professor King has published a valuable memoir on the Permian fossils of England, in the Proceedings of the Palæontographical Society, in which the following Table is given (in descending order) of the Permian system of the North of England, as compared with that of Thuringia:—
At the base of the system lies a band of lower sandstone ([No. 6]) of various colours, separating the Magnesian Limestone from the coal in Yorkshire and Durham; sometimes associated with red marl and gypsum, but with the same obscure relations in all these beds which usually attend the close of one series and the commencement of another; the imbedded plants being, in some cases, stated to be identical with those of the Carboniferous series. In Thuringia the Rothliegende, or red-lyer, a great deposit of red sandstone and conglomerate, associated with porphyry, basaltic trap, and amygdaloid, lies at the base of the system. Among the fossils of this age are the silicified trunks of Tree-ferns (Psaronius), the bark of which is surrounded by dense masses of air-roots, which often double or quadruple the diameter of the original stem; in this respect bearing a strong resemblance to the living arborescent ferns of New Zealand.
The marl-slate ([No. 5]) consists of hard calcareous shales, marl-slates, and thin-bedded limestone, the whole nearly thirty feet thick in Durham, and yielding many fine specimens of Ganoid and Placoid fishes—Palæoniscus, Pygopterus, Cœlacanthus, and Platysomus—genera which all belong to the Carboniferous system, and which Professor King thinks probably lived at no great distance from the shore; but the Permian species of the marl-slate of England are identical with those of the copper-slate of Thuringia. Agassiz was the first to point out a remarkable peculiarity in the forms of the fishes which lived before and after this period. In most living fishes the trunk seems to terminate in the middle of the root of the tail, whose free margin is “homocercal” (even-tail), that is, either rounded, or, if forked, divided into two equal lobes. In Palæoniscus, and most Palæozoic fishes, the axis of the body is continued into the upper lobe of the tail, which is thus rendered unsymmetrical, as in the living sharks and sturgeons. The latter form, which Agassiz termed “heterocercal” (unequal-tail) is only in a very general way distinctive of Palæozoic fishes, since this asymmetry exists, though in a minor degree, in many living genera besides those just mentioned. The compact limestone ([No. 4]) is rich in Polyzoa. The fossiliferous limestone ([No. 3]), Mr. King considers, is a deep-water formation, from the numerous Polyzoa which it contains. One of these, Fenestella retiformis, found in the Permian rocks of England and Germany, sometimes measures eight inches in width.
Many species of Mollusca, and especially Brachiopoda, appear in the Permian seas of this age, Spirifera and Producta being the most characteristic.