In the swamps and ditches of England there grows a plant called the horse-tail (equisetum), having a succulent, erect, jointed stem, with slender leaves, and a scaly catkin at the top. A second large section of the plants of the carboniferous era were of this kind (equisetaceæ), but, like the fern, reaching the magnitudes of trees. While existing equiseta rarely exceed three feet in height, and the stems are generally under half an inch in diameter, their kindred, entombed in the coal beds, seem to have been generally fourteen or fifteen feet high, with stems from six inches to a foot in thickness. Arborescent plants of this family, like the arborescent ferns, now grow only in tropical countries, and their being found in the coal beds in all latitudes is consequently held as an additional proof, that at this era a warm climate was extended much farther to the north than at present. It is to be remarked that plants of this kind (forming two genera, the most abundant of which is the calamites) are only represented on the present surface by plants of the same family: the species which flourished at this era gradually lessen in number as we advance upwards in the series of rocks, and disappear before we arrive at the tertiary formation.
The club-moss family (lycopodiaceæ) are other plants of the present surface, usually seen in a lowly and creeping form in temperate latitudes, but presenting species which rise to a greater magnitude within the tropics. Many specimens of this family are found in the coal beds; it is thought they have contributed more to the substance of the coal than any other family. But, like the ferns and equisetaceæ, they rise to a prodigious magnitude. The lepidodendra (so the fossil genus is called) have probably been from sixty-five to eighty feet in height, having at their base a diameter of about three feet, while their leaves measured twenty inches in length. In the forests of the coal era, the lepidodendra would enjoy the rank of firs in our forests, affording shade to the only less stately ferns and calamites. The internal structure of the stem, and the character of the seed-vessels, shew them to have been a link between single-lobed and double-lobed plants, a fact worthy of note, as it favours the idea that, in vegetable, as well as animal creation, a progress has been observed, in conformity with advancing conditions. It is also curious to find a missing link of so much importance in a genus of plants which has long ceased to have a living place upon earth.
The other leading plants of the coal era are without representatives on the present surface, and their characters are in general less clearly ascertained. Amongst the most remarkable are—the sigillaria, of which large stems are very abundant, shewing that the interior has been soft, and the exterior fluted with separate leaves inserted in vertical rows along the flutings—and the stigmaria, plants apparently calculated to flourish in marshes or pools, having a short, thick, fleshy stem, with a dome-shaped top, from which sprung branches of from twenty to thirty feet long. Amongst monocotyledons were some palms, (flabellaria and næggerathia,) besides a few not distinctly assignable to any class.
The dicotyledons of the coal are comparatively few, though on the present surface they are the most numerous sub-class. Besides some of doubtful affinity, (annularia, asterophyllites, &c.,) there were a few of the pine family, which seem to have been the highest class of trees of this era, and are only as yet found in isolated cases, and in sandstone beds. The first discovered lay in the Craigleith quarry, near Edinburgh, and consisted of a stem about two feet thick, and forty-seven feet in length. Others have since been found, both in the same situation, and at Newcastle. Leaves and fruit being wanting, an ingenious mode of detecting the nature of these trees was hit upon by Mr. Witham of Lartington. Taking thin polished cross slices of the stem, and subjecting them to the microscope, he detected the structure of the wood to be that of a cone-bearing tree, by the presence of certain “reticulations” which distinguish that family, in addition to the usual radiating and concentric lines. That particular tree was concluded to be an araucaria, a species now found in Norfolk Island, in the South Sea, and in a few other remote situations. The coniferæ of this era form the dawn of dicotyledenous trees, of which they may be said to be the simplest type, and to which, it has already been noticed, the lepidodendra are a link from the monocotyledons. The concentric rings of the Craigleith and other coniferæ of this era have been mentioned. It is interesting to find in these a record of the changing seasons of those early ages, when as yet there were no human beings to observe time or tide. They are clearly traced; but it is observed that they are more slightly marked than is the case with their family at the present day, as if the changes of temperature had been within a narrower range.
Such was the vegetation of the carbonigenous era, composed of forms at the bottom of the botanical scale, flowerless, fruitless, but luxuriant and abundant beyond what the most favoured spots on earth can now shew. The rigidity of the leaves of its plants, and the absence of fleshy fruits and farinaceous seeds, unfitted it to afford nutriment to animals; and, monotonous in its forms, and destitute of brilliant colouring, its sward probably unenlivened by any of the smaller flowering herbs, its shades uncheered by the hum of insects, or the music of birds, it must have been but a sombre scene to a human visitant. But neither man nor any other animals were then in existence to look for such uses or such beauties in this vegetation. It was serving other and equally important ends, clearing (probably) the atmosphere of matter noxious to animal life, and storing up mineral masses which were in long subsequent ages to prove of the greatest service to the human race, even to the extent of favouring the progress of its civilization.
The animal remains of this era are not numerous, in comparison with those which go before, or those which come after. The mountain limestone, indeed, deposited at the commencement of it, abounds unusually in polypiaria and crinoidea; but when we ascend to the coal-beds themselves, the case is altered, and these marine remains altogether disappear. We have then only a limited variety of conchifers and shell mollusks, with fragments of a few species of fishes, and these are rarely or never found in the coal seams, but in the shales alternating with them. Some of the fishes are of a sauroid character, that is, partake of the nature of the lizard, a genus of the reptilia, a land class of animals, so that we may be said here to have the first approach to a kind of animals calculated to breathe the atmosphere. Such is the Megalichthys Hibbertii, found by Dr. Hibbert Ware, in a limestone bed of fresh-water origin, underneath the coal at Burdiehouse, near Edinburgh. Others of the same kind have been found in the coal measures in Yorkshire, and in the low coal shales at Manchester. This is no more than might be expected, as collections of fresh water now existed, and it is presumable that they would be peopled. The chief other fishes of the coal era are named palæothrissum, palæoniscus, diperdus.
Coal strata are nearly confined to the group termed the carboniferous formation. Thin beds are not unknown afterwards, but they occur only as a rare exception. It is therefore thought that the most important of the conditions which allowed of so abundant a terrestrial vegetation, had ceased about the time when this formation was closed. The high temperature was not one of the conditions which terminated, for there are evidences of it afterwards; but probably the superabundance of carbonic acid gas supposed to have existed during this era was expended before its close. There can be little doubt that the infusion of a large dose of this gas into the atmosphere at the present day would be attended by precisely the same circumstances as in the time of the carboniferous formation. Land animal life would not have a place on earth; vegetation would be enormous; and coal strata would be formed from the vast accumulations of woody matter, which would gather in every sea, near the mouths of great rivers. On the exhaustion of the superabundance of carbonic acid gas, the coal formation would cease, and the earth might again become a suitable theatre of being for land animals.
The termination of the carboniferous formation is marked by symptoms of volcanic violence, which some geologists have considered to denote the close of one system of things and the beginning of another. Coal beds generally lie in basins, as if following the curve of the bottom of seas. But there is no such basin which is not broken up into pieces, some of which have been tossed up on edge, others allowed to sink, causing the ends of strata to be in some instances many yards, and in a few several hundred feet, removed from the corresponding ends of neighbouring fragments. These are held to be results of volcanic movements below, the operation of which is further seen in numerous upbursts and intrusions of volcanic rock (trap). That these disturbances took place about the close of the formation, and not later, is shewn in the fact of the next higher group of strata being comparatively undisturbed. Other symptoms of this time of violence are seen in the beds of conglomerate which occur amongst the first strata above the coal. These, as usual, consist of fragments of the elder rocks, more or less worn from being tumbled about in agitated water, and laid down in a mud paste, afterwards hardened. Volcanic disturbances break up the rocks; the pieces are worn in seas; and a deposit of conglomerate is the consequence. Of porphyry, there are some such pieces in the conglomerate of Devonshire, three or four tons in weight. It is to be admitted for strict truth that, in some parts of Europe, the carboniferous formation is followed by superior deposits, without the appearance of such disturbances between their respective periods; but apparently this case belongs to the class of exceptions already noticed. [93] That disturbance was general, is supported by the further and important fact of the destruction of many forms of organic being previously flourishing, particularly of the vegetable kingdom.
ERA OF THE NEW RED SANDSTONE.
TERRESTRIAL ZOOLOGY COMMENCES
WITH REPTILES.
FIRST TRACES OF BIRDS.
The next volume of the rock series refers to an era distinguished by an event of no less importance than the commencement of land animals. The New Red Sandstone System is subdivided into groups, some of which are wanting in some places; they are pretty fully developed in the north of England, in the following ascending order:—1. Lower red sandstone; 2. Magnesian limestone; 3. Red and white sandstones and conglomerate; 4. Variegated marls. Between the third and fourth there is, in Germany, another group, called the Muschelkalk, a word expressing a limestone full of shells.