The first group, containing the conglomerates already adverted to, seems to have been produced during the time of disturbance which occurred so generally after the carbonigenous era. This new era is distinguished by a paucity of organic remains, as might partly be expected from the appearances of disturbance, and the red tint of the rocks, the latter being communicated by a solution of oxide of iron, a substance unfavourable to animal life.

The second group is a limestone with an infusion of magnesia. It is developed less generally than some others, but occurs conspicuously in England and Germany. Its place, above the red sandstone, shews the recurrence of circumstances favourable to animal life, and we accordingly find in it not only zoophytes, conchifera, and a few tribes of fish, but some faint traces of land plants, and a new and startling appearance—a reptile of saurian (lizard) character, analogous to the now existing family called monitors. Remains of this creature are found in cupriferous (copper-bearing) slate connected with the mountain limestone, at Mansfield and Glucksbrunn, in Germany, which may be taken as evidence that dry land existed in that age near those places. The magnesia limestone is also remarkable as the last rock in which appears the leptæna, or producta, a conchifer of numerous species which makes a conspicuous appearance in all previous seas. It is likewise to be observed, that the fishes of this age, to the genera of which the names palæoniscus, catopterus, platysomus, &c., have been applied, vanish, and henceforth appear no more.

The third group, chiefly sandstones, variously coloured according to the amount and nature of the metallic oxide infused into them, shews a recurrence of agitation, and a consequent diminution of the amount of animal life. In the upper part, however, of this group, there are abundant symptoms of a revival of proper conditions for such life. There are marl beds, the origin of which substance in decomposed shells is obvious; and in Germany, though not in England, here occurs the muschelkalk, containing numerous organic remains, (generally different from those of the magnesian limestone,) and noted for the specimens of land animals, which it is the first to present in any considerable abundance to our notice.

These animals are of the vertebrate sub-kingdom, but of its lowest class next after fishes,—namely, reptiles,—a portion of the terrestrial tribes whose imperfect respiratory system perhaps fitted them for enduring an atmosphere not yet quite suitable for birds or mammifers. [97] The specimens found in the muschelkalk are allied to the crocodile and lizard tribes of the present day, but in the latter instance are upon a scale of magnitude as much superior to present forms as the lepidodendron of the coal era was superior to the dwarf club-mosses of our time. These saurians also combine some peculiarities of structure of a most extraordinary character.

The animal to which the name ichthyosaurus has been given, was as long as a young whale, and it was fitted for living in the water, though breathing the atmosphere. It had the vertebral column and general bodily form of a fish, but to that were added the head and breast-bone of a lizard, and the paddles of the whale tribes. The beak, moreover, was that of a porpoise, and the teeth were those of a crocodile. It must have been a most destructive creature to the fish of those early seas.

The plesiosaurus was of similar bulk, with a turtle-like body and paddles, shewing that the sea was its element, but with a long serpent-like neck, terminating in a saurian head, calculated to reach prey at a considerable distance. These two animals, of which many varieties have been discovered, constituting distinct species, are supposed to have lived in the shallow borders of the seas of this and subsequent formations, devouring immense quantities of the finny tribes. It was at first thought that no creatures approaching them in character now inhabit the earth; but latterly Mr. Darwin has discovered, in the reptile-peopled Galapagos Islands, in the South Sea, a marine saurian from three to four feet long.

The megalosaurus was an enormous lizard—a land creature, also carnivorous. The pterodactyle was another lizard, but furnished with wings to pursue its prey in the air, and varying in size between a cormorant and a snipe. Crocodiles abounded, and some of these were herbivorous. Such was the iguanodon, a creature of the character of the iguana of the Ganges, but reaching a hundred feet in length, or twenty times that of its modern representative.

There were also numerous tortoises, some of them reaching a great size; and Professor Owen has found in Warwickshire some remains of an animal of the batrachian order, [99] to which, from the peculiar form of the teeth, he has given the name of labyrinthidon. Thus, three of Cuvier’s four orders of reptilia (sauria, chelonia, and batrachia) are represented in this formation, the serpent order (ophidia) being alone wanting.

The variegated marl beds which constitute the uppermost group of the formation, present two additional genera of huge saurians,—the phytosaurus and mastodonsaurus.

It is in the upper beds of the red sandstone that beds of salt first occur. These are sometimes of such thickness, that the mine from which the material has been excavated looks like a lofty church. We see in the present world no circumstances calculated to produce the formation of a bed of rock salt; yet it is not difficult to understand how such strata were formed in an age marked by ultra-tropical heat and frequent volcanic disturbances. An estuary, cut off by an upthrow of trap, or a change of level, and left to dry up under the heat of the sun, would quickly become the bed of a dense layer of rock salt. A second shift of level, or some other volcanic disturbance, connecting it again with the sea, would expose this stratum to being covered over with a layer of sand or mud, destined in time to form the next stratum of rock above it.