In the second stage, the mother substance is capable of giving off two smaller cleavage products, Y and B. Y is split off by the long waves of light, and B by the short waves, and the vital response to Y is the sensation of yellow, that to B the sensation of blue. But suppose that, chemically, Y + B = W: then, if Y and B are both split off at the same time in the same cone, they immediately unite into W, and the resulting sensation is white, and neither yellow nor blue.
Fig. 38.--The cleavage products, in the three stages of the color sense. The "mother substance" is not represented in the diagram, but only the cleavage products which, according to the Ladd-Franklin theory, are the direct stimuli for the color sensations. (Figure text: 1--white, 2--yellow blue, 3--red green blue)
Similarly, in the third stage, the mother substance is capable of giving off three cleavage products, R, G and B; and there are three corresponding vital responses, the sensations of red, green and blue. But, chemically, R + G = Y; and therefore, if R and G are split off at the same time, they unite chemically into Y and give the sensation of yellow. If R, G and B are all split off at the same time, they unite chemically as follows: R + G = Y, and Y + B = W; and therefore the resulting sensation is that of white.
This theory of cleavage products is in good general agreement with chemical principles, and it does justice to all the facts of color vision, as detailed in the preceding pages. It should be added that "for black, the theory supposes that, [{224}] in the interest of a continuous field of view, objects which reflect no light at all upon the retina have correlated with them a definite non-light sensation--that of black." [Footnote: Quotation from Dr. Ladd-Franklin.]
Adaptation
Sensory adaptation is a change that occurs in other senses also, but it is so much more important in the sense of sight than elsewhere that it may best be considered here. The stimulus continues, the sensation ceases or diminishes--that is the most striking form of sensory adaptation. Continued action of the same stimulus puts the sense into such a condition that it responds differently from at first, and usually more weakly. It is much like fatigue, but it often is more positive and beneficial than fatigue.
The sense of smell is very subject to adaptation. On first entering a room you clearly sense an odor that you can no longer get after staying there for some time. This adaptation to one odor does not prevent your sensing quite different odors. Taste shows less adaptation than smell, but all are familiar with the decline in sweet sensation that comes with continued eating of sweets.
All of the cutaneous senses except that for pain are much subject to adaptation. Continued steady pressure gives a sensation that declines rapidly and after a time ceases altogether. The temperature sense is usually adapted to the temperature of the skin, which therefore feels neither warm nor cool. If the temperature of the skin is raised from its usual level of about 70 degrees Fahrenheit to 80 or 86, this temperature at first gives the sensation of warmth, but after a time it gives no temperature sensation at all; the warmth sense has become adapted to the temperature of 80 degrees; and now a temperature of 70 will give the sensation of cool. [{225}] Hold one hand in water at 80 and the other in water at 66, and when both have become adapted to these respective temperatures, plunge them together into water at 70; and you will find this last to feel cool to the warm-adapted hand and warm to the cool-adapted. There are limits to this power of adaptation.