The Antiquity of Man

by Sir Charles Lyell

Mr. Darwin's Theory of the Origin of Species by Natural Selection.
Memoir by Mr. Wallace.
Manner in which favoured Races prevail in the Struggle for Existence.
Formation of new Races by breeding.
Hypotheses of definite and indefinite Modifiability equally arbitrary.
Competition and Extinction of Races.
Progression not a necessary Accompaniment of Variation.
Distinct Classes of Phenomena which Natural Selection explains.
Unity of Type, Rudimentary Organs, Geographical Distribution,
Relation of the extinct to the living Fauna and Flora, and
mutual Relations of successive Groups of Fossil Forms.
Light thrown on Embryological Development by Natural Selection.
Why large Genera have more variable Species than small ones.
Dr. Hooker on the Evidence afforded by the Vegetable Kingdom
in favour of Creation by Variation.
Steenstrup on alternation of Generations.
How far the Doctrine of Independent Creation is opposed to the
Laws now governing the Migration of Species.

For many years after the promulgation of Lamarck's doctrine of progressive development, geologists were much occupied with the question whether the past changes in the animate and inanimate world were brought about by sudden and paroxysmal action, or gradually and continuously, by causes differing neither in kind nor degree from those now in operation.

The anonymous author of "The Vestiges of Creation" published in 1844 a treatise, written in a clear and attractive style, which made the English public familiar with the leading views of Lamarck on transmutation and progression, but brought no new facts or original line of argument to support those views, or to combat the principal objections which the scientific world entertained against them.

No decided step in this direction was made until the publication in 1858 of two papers, one by Mr. Darwin and another by Mr. Wallace, followed in 1859 by Mr. Darwin's celebrated work on "The Origin of Species by Means of Natural Selection; or, the Preservation of favoured Races in the Struggle for Life." The author of this treatise had for twenty previous years strongly inclined to believe that variation and the ordinary laws of reproduction were among the secondary causes always employed by the Author of nature, in the introduction from time to time of new species into the world, and he had devoted himself patiently to the collecting of facts and making of experiments in zoology and botany, with a view of testing the soundness of the theory of transmutation. Part of the manuscript of his projected work was read to Dr. Hooker as early as 1844 and some of the principal results were communicated to me on several occasions. [Note 40] Dr. Hooker and I had repeatedly urged him to publish without delay, but in vain, as he was always unwilling to interrupt the course of his investigations; until at length Mr. Alfred R. Wallace, who had been engaged for years in collecting and studying the animals of the East Indian archipelago, thought out independently for himself one of the most novel and important of Mr. Darwin's theories. This he embodied in an essay "On the Tendency of Varieties to depart indefinitely from the original Type." It was written at Ternate in February 1858, and sent to Mr. Darwin with a request that it might be shown to me if thought sufficiently novel and interesting. Dr. Hooker and I were of opinion that it should be immediately printed, and we succeeded in persuading Mr. Darwin to allow one of the manuscript chapters of his "Origin of Species," entitled "On the Tendency of Species to form Varieties, and on the Perpetuation of Species and Varieties by natural Means of Selection," to appear at the same time.*

(* See "Proceedings of the Linnaean Society" 1858.)

By reference to these memoirs it will be seen that both writers begin by applying to the animal and vegetable worlds the Malthusian doctrine of population, or its tendency to increase in a geometrical ratio, while food can only be made to augment even locally in an arithmetical one. There being therefore no room or means of subsistence for a large proportion of the plants and animals which are born into the world, a great number must annually perish. Hence there is a constant struggle for existence among the individuals which represent each species and the vast majority can never reach the adult state, to say nothing of the multitudes of ova and seeds which are never hatched or allowed to germinate. Of birds it is estimated that the number of those which die every year equals the aggregate number by which the species to which they respectively belong is on the average permanently represented.

The trial of strength which must decide what individuals are to survive and what to succumb occurs in the season when the means of subsistence are fewest, or enemies most numerous, or when the individuals are enfeebled by climate or other causes; and it is then that those varieties which have any, even the slightest, advantage over others come off victorious. They may often owe their safety to what would seem to a casual observer a trifling difference, such as a darker or lighter shade of colour rendering them less visible to a species which preys upon them, or sometimes to attributes more obviously advantageous, such as greater cunning or superior powers of flight or swiftness of foot. These peculiar qualities and faculties, bodily and instinctive, may enable them to outlive their less favoured rivals, and being transmitted by the force of inheritance to their offspring will constitute new races, or what Mr. Darwin calls "incipient species." If one variety, being in other respects just equal to its competitors, happens to be more prolific, some of its offspring will stand a greater chance of being among those which will escape destruction, and their descendants, being in like manner very fertile, will continue to multiply at the expense of all less prolific varieties.

As breeders of domestic animals, when they choose certain varieties in preference to others to breed from, speak technically of their method as that of "selecting," Mr. Darwin calls the combination of natural causes, which may enable certain varieties of wild animals or plants to prevail over others of the same species, "natural selection."

A breeder finds that a new race of cattle with short horns or without horns may be formed in the course of several generations by choosing varieties having the most stunted horns as his stock from which to breed; so nature, by altering in the course of ages, the conditions of life, the geographical features of a country, its climate, the associated plants and animals, and consequently the food and enemies of a species and its mode of life, may be said, by this means to select certain varieties best adapted for the new state of things. Such new races may often supplant the original type from which they have diverged, although that type may have been perpetuated without modification for countless anterior ages in the same region, so long as it was in harmony with the surrounding conditions then prevailing.

Lamarck, when speculating on the origin of the long neck of the giraffe, imagined that quadruped to have stretched himself up in order to reach the boughs of lofty trees, until by continued efforts and longing to reach higher he obtained an elongated neck. Mr. Darwin and Mr. Wallace simply suppose that, in a season of scarcity, a longer-necked variety, having the advantage in this respect over most of the herd, as being able to browse on foliage out of their reach, survived them and transmitted its peculiarity of cervical conformation to its successors.