(* Wollaston, "On the Variation of Species" etc. London
1856.)
(** "Transactions of Northern Entomological Society" 1862.)
The reader will remember that at the commencement of the Glacial period there was scarcely any appreciable difference between the molluscous fauna and that now living. When therefore the events of the Glacial period, as described in the earlier part of this volume, are duly pondered on, and when we reflect that in the Upper Miocene period the living species of mollusca constitute only one-third of the whole fauna, we see clearly by how high a figure we must multiply the time in order to express the distance between the Miocene period and our own days.
SPECIES OF MAMMALIA RECENT AND FOSSIL—PROBOSCIDIANS.
But it may perhaps be said that the mammalia afford more conspicuous examples than do the mollusca, insects, or plants of the wide gaps which separate species and genera, and that if in this higher class such a multitude of transitional forms had ever existed as would be required to unite the Tertiary and Recent species into one series or net-work of allied or transitional forms, they could not so entirely have escaped observation whether in the fossil or living fauna. A zoologist who entertains such an opinion would do well to devote himself to the study of some one genus of mammalia, such as the elephant, rhinoceros, hippopotamus, bear, horse, ox, or deer; and after collecting all the materials he can get together respecting the extinct and Recent species, decide for himself whether the present state of science justifies his assuming that the chain could never have been continuous, the number of the missing links being so great.
Among the extinct species formerly contemporary with man, no fossil quadruped has so often been alluded to in this work as the mammoth, Elephas primigenius. From a monograph on the proboscidians by Dr. Falconer, it appears that this species represents one extreme of a type of which the Pliocene Mastodon borsoni represents the other. Between these extremes there are already enumerated by Dr. Falconer no less than twenty-six species, some of them ranging as far back in time as the Miocene period, others still living, like the Indian and African forms. Two of these species, however, he has always considered as doubtful, Stegodon ganesa, probably a mere variety of one of the others, and Elephas priscus of Goldfuss, founded partly on specimens of the African elephant, assumed by mistake to be fossil, and partly on some aberrant forms of E. antiquus.
The first effect of the intercalation of so many intermediate forms between the two most divergent types, has been to break down almost entirely the generic distinction between Mastodon and Elephas. Dr. Falconer, indeed, observes that Stegodon (one of several subgenera which he has founded) constitutes an intermediate group, from which the other species diverge through their dental characters, on the one side into the mastodons, and on the other into the Elephants.*
(* "Quarterly Journal of the Geological Society" volume 13
1857 page 314.)
The next result is to diminish the distance between the several members of each of these groups.
Dr. Falconer has discovered that no less than four species of elephant were formerly confounded together under the title of Elephas primigenius, whence its supposed ubiquity in Pleistocene times, or its wide range over half the habitable globe. But even when this form has been thus restricted in its specific characters, it has still its geographical varieties; for the mammoth's teeth brought from America may in most instances, according to Dr. Falconer, be distinguished from those proper to Europe. On this American variety Dr. Leidy has conferred the name of E. americanus. Another race of the same mammoth (as determined by Dr. Falconer) existed, as we have seen, before the Glacial period, or at the time when the buried forest of Cromer and the Norfolk cliffs was deposited; and the Swiss geologists have lately found remains of the mammoth in their country, both in pre-glacial and post-glacial formations.
Since the publication of Dr. Falconer's monograph, two other species of elephant, F. mirificus, Leidy, and F. imperator, have been obtained from the Pliocene formations of the Niobrara Valley in Nebraska, one of which, however, may possibly be found hereafter to be the same as E. columbi, Falc. A remarkable dwarf species also (Elephas melitensis) has been discovered, belonging, like the existing E. africanus, to the group Loxodon. This species has been established by Dr. Falconer on remains found by Captain Spratt R.N. in a cave in Malta.*