The Antiquity of Man

by Sir Charles Lyell

(* "Proceedings of the Geological Society" London 1862.)

How much the difficulty of discriminating between the fossil representatives of this genus may hereafter augment, when all the species with their respective geographical varieties are known, may be inferred from the following fact—Professor H. Schlegel, in a recently published memoir, endeavours to show that the living elephant of Sumatra agrees with that of Ceylon, but is a distinct species from that of Continental India, being distinguishable by the number of its dorsal vertebrae and ribs, the form of its teeth, and other characteristics.*

(* Schlegel, "Natural History Review" Number 5 1862 page
72.)

Dr. Falconer, on the other hand, considers these two living species as mere geographical varieties, the characters referred to not being constant, as he has ascertained, on comparing different individuals of E. indicus in different parts of Bengal in which the ribs vary from nineteen to twenty, and different varieties of E. africanus in which they vary from twenty to twenty-one.

An inquiry into the various species of the genus Rhinoceros, recent and fossil, has led Dr. Falconer to analogous results, as might be inferred from what was said in Chapter 10, and as a forthcoming memoir by the same writer will soon more fully demonstrate.

Among the fossils brought in 1858 by Mr. Hayden from the Niobrara Valley, Dr. Leidy describes a rhinoceros so like the Asiatic species, R. indicus, that he at first referred it to the same, and, what is most singular, he remarks generally of the Pliocene fauna of that part of North America that it is far more related in character to the Pleistocene and Recent fauna of Europe than to that now inhabiting the American continent.

It seems indeed more and more evident that when we speculate in future on the pedigree of any extinct quadruped which abounds in the drift or caverns of Europe, we shall have to look to North and South America as a principal source of information. Thirty years ago, if we had been searching for fossil types which might fill up a gap between two species or genera of the horse tribe (or great family of the Solipedes), we might have thought it sufficient to have got together as ample materials as we could obtain from the continents of Europe, Africa, and Asia. We might have presumed that as no living representative of the equine family, whether horse, ass, zebra, or quagga, had been furnished by North or South America when those regions were first explored by Europeans, a search in the transatlantic world for fossil species might be dispensed with. But how different is the prospect now opening before us! Mr. Darwin first detected the remains of a fossil horse during his visit to South America, since which two other species have been met with on the same continent, while in North America, in the valley of the Nebraska alone, Mr. Hayden, besides a species not distinguishable from the domestic horse, has obtained, according to Dr. Leidy, representatives of five other fossil genera of Solipedes. These he names, Hipparion, Protohippus, Merychippus, Hypohippus, and Parahippus. On the whole, no less than twelve equine species, belonging to seven genera (including the Miocene Anchitherium of Nebraska), being already detected in the Tertiary and Post-Tertiary formations of the United States.*

(* "Proceedings of the Academy of Natural Science"
Philadelphia for 1858 page 89.)

Professors Unger* and Heer** have advocated, on botanical grounds, the former existence of an Atlantic continent during some part of the Tertiary period, as affording the only plausible explanation that can be imagined, of the analogy between the Miocene flora of Central Europe and the existing flora of Eastern America. Professor Oliver, on the other hand, after showing how many of the American types found fossil in Europe are common to Japan, inclines to the theory, first advanced by Dr. Asa Gray, that the migration of species, to which the community of types in the eastern states of North America and the Miocene flora of Europe is due, took place when there was an overland communication from America to eastern Asia between the fiftieth and sixtieth parallels of latitude, or south of Behring Straits, following the direction of the Aleutian islands.*** By this course they may have made their way, at any epoch, Miocene, Pliocene, or Pleistocene, antecedently to the glacial epoch, to Mongolia, on the east coast of northern Asia.

(* "Die versunkene Insel Atlantis.")
(** "Flora tertiaria Helvetiae.")
(*** Oliver, Lecture at the Royal Institution, March 7, 1862.)