Glaciation of Wales and England.—The mountains of North Wales were recognised, in 1842, by Dr. Buckland, as having been an independent centre of the dispersion of erratics—great glaciers, long since extinct, having radiated from the Snowdonian heights in Carnarvonshire, through seven principal valleys towards as many points of the compass, carrying with them large stony fragments, and grooving the subjacent rocks in as many directions.

Besides this evidence of land-glaciers, Mr. Trimmer had previously, in 1831, detected the signs of a great submergence in Wales in the Post-pliocene period. He had observed stratified drift, from which he obtained about a dozen species of marine shells, near the summit of Moel Tryfaen, a hill 1400 feet high, on the south side of the Menai Straits. I had an opportunity of examining in the summer of 1863, together with the Reverend W. S. Symonds, a long and deep cutting made through this drift by the Alexandra Mining Company in search of slates. At the top of the hill above-mentioned we saw a stratified mass of incoherent sand and gravel 35 feet thick, from which no less than 54 species of mollusca, besides three characteristic arctic varieties—in all 57 forms—have been obtained by Mr. Darbishire. They belong without exception to species still living in British or more northern seas; eleven of them being exclusively arctic, four common to the arctic and British seas, and a large proportion of the remainder having a northward range, or, if found at all in the southern seas of Britain, being comparatively less abundant. In the lowest beds of the drift were large heavy boulders of far-transported rocks, glacially polished and scratched on more than one side. Underneath the whole we saw the edges of vertical slates exposed to view, which here, like the rocks in other parts of Wales, both at greater and less elevations, exhibit beneath the drift unequivocal marks of prolonged glaciation. The whole deposit has much the appearance of an accumulation in shallow water or on a beach, and it probably acquired its thickness during the gradual subsidence of the coast—an hypothesis which would require us to ascribe to it a high antiquity, since we must allow time, first for its sinking, and then for its re-elevation.

The height reached by these fossil shells on Moel Tryfaen is no less than 1300 feet—a most important fact when we consider how very few instances we have on record beyond the limits of Wales, whether in Europe or North America, of marine shells having been found in glacial drift at half the height above indicated. A marine molluscous fauna, however, agreeing in character with that of Moel Tryfaen, and comprising as many species, has been found in drift at Macclesfield and other places in central England, sometimes reaching an elevation of 1200 feet.

Professor Ramsay[[5]] estimated the probable amount of submergence during some part of the glacial period at about 2300 feet; for he was unable to distinguish the superficial sands and gravel which reached that high elevation from the drift which, at Moel Tryfaen and at lower points, contains shells of living species. The evidence of the marine origin of the highest drift is no doubt inconclusive in the absence of shells, so great is the resemblance of the gravel and sand of a sea beach and of a river’s bed, when organic remains are wanting; but, on the other hand, when we consider the general rarity of shells in drift which we know to be of marine origin, we cannot suppose that, in the shelly sands of Moel Tryfaen, we have hit upon the exact uppermost limit of marine deposition, or, in other words, a precise measure of the submergence of the land beneath the sea since the glacial period.

We are gradually obtaining proofs of the larger part of England, north of a line drawn from the mouth of the Thames to the Bristol Channel, having been under the sea and traversed by floating ice since the commencement of the glacial epoch. Among recent observations illustrative of this point, I may allude to the discovery, by Mr. J. F. Bateman, near Blackpool, in Lancashire, fifty miles from the sea, and at the height of 568 feet above its level, of till containing rounded and angular stones and marine shells, such as Turritella communis, Purpura lapillus, Cardium edule, and others, among which Trophon clathratum (=Fusus Bamffius), though still surviving in North British seas, indicates a cold climate.

Erratics near Chichester.—The most southern memorials of ice-action and of a Post-pliocene fauna in Great Britain is on the coast of the county of Sussex, about 25 miles west of Brighton, and 15 south of Chichester. A marine deposit exposed between high and low tide occurs on both sides of the promontory called Selsea Bill, in which Mr. Godwin-Austen found thirty-eight species of shells, and the number has since been raised to seventy.

This assemblage is interesting because on the whole, while all the species are recent, they have a somewhat more southern aspect than those of the present British Channel. It is true that about forty of them range from British to high northern latitudes; but several of them, as, for example, Lutraria rugosa and Pecten polymorphous, which are abundant, are not known at present to range farther north than the coast of Portugal, and seem to indicate a warmer temperature than now prevails on the coast where we find them fossil. What renders this curious is the fact that the sandy loam in which they occur is overlaid by yellow clayey gravel with large erratic blocks which must have been drifted into their present position by ice when the climate had become much colder. These transported fragments of granite, syenite, and greenstone, as well as of Devonian and Silurian rocks, may have come from the coast of Normandy and Brittany, and are many of them of such large size that we must suppose them to have been drifted into their present site by coast-ice. I measured one of granite, at Pagham, 21 feet in circumference. In the gravel of this drift with erratics are a few littoral shells of living species, indicating an ancient coast-line.

Glacial Formations of North America.—In the western hemisphere, both in Canada and as far south as the 40th and even 38th parallel of latitude in the United States, we meet with a repetition of all the peculiarities which distinguish the European boulder formation. Fragments of rock have travelled for great distances, especially from north to south: the surface of the subjacent rock is smoothed, striated, and fluted; unstratified mud or till containing boulders is associated with strata of loam, sand, and clay, usually devoid of fossils. Where shells are present, they are of species still living in northern seas, and not a few of them identical with those belonging to European drift, including most of those already given in Figs. 107 to 112, p. 176. The fauna also of the glacial epoch in North America is less rich in species than that now inhabiting the adjacent sea, whether in the Gulf of St. Lawrence, or off the shores of Maine, or in the Bay of Massachusetts.

The extension on the American continent of the range of erratics during the Post-pliocene period to lower latitudes than they reached in Europe, agrees well with the present southward deflection of the isothermal lines, or rather the lines of equal winter temperature. It seems that formerly, as now, a more extreme climate and a more abundant supply of ice prevailed on the western side of the Atlantic. Another resemblance between the distribution of the drift fossils in Europe and North America has yet to be pointed out. In Canada and the United States, as in Europe, the marine shells are generally confined to very moderate elevations above the sea (between 100 and 700 feet), while the erratic blocks and the grooved and polished surfaces of rock extend to elevations of several thousand feet.

I have already mentioned that in Europe several quadrupeds of living, as well as extinct, species were common to pre-glacial and post-glacial times. In like manner there is reason to suppose that in North America much of the ancient mammalian fauna, together with nearly all the invertebrata, lived through the ages of intense cold. That in the United States the Mastodon giganteus was very abundant after the drift period, is evident from the fact that entire skeletons of this animal are met with in bogs and lacustrine deposits occupying hollows in the glacial drift. They sometimes occur in the bottom even of small ponds recently drained by the agriculturist for the sake of the shell-marl. In 1845 no less than six skeletons of the same species of Mastodon were found in Warren county, New Jersey, six feet below the surface, by a farmer who was digging out the rich mud from a small pond which he had drained. Five of these skeletons were lying together, and a large part of the bones crumbled to pieces as soon as they were exposed to the air.