It would be rash, however, to infer from such data that these quadrupeds were mired in modern times, unless we use that term strictly in a geological sense. I have shown that there is a fluviatile deposit in the valley of the Niagara, containing shells of the genera Melania, Lymnea, Planorbis, Velvata, Cyclaz, Unio, Helix, etc., all of recent species, from which the bones of the great Mastodon have been taken in a very perfect state. Yet the whole excavation of the ravine, for many miles below the Falls, has been slowly effected since that fluviatile deposit was thrown down. Other extinct animals accompany the Mastodon giganteus in the post-glacial deposits of the United States, and this, taken with the fact that so few of the mollusca, even of the commencement of the cold period, differ from species now living is important, as refuting the hypothesis, for which some have contended, that the intensity of the glacial cold annihilated all the species in temperate and arctic latitudes.

Connection of the Predominance of Lakes with Glacial Action.—It was first pointed out by Professor Ramsay in 1862, that lakes are exceedingly numerous in those countries where erratics, striated blocks, and other signs of ice-action abound; and that they are comparatively rare in tropical and sub-tropical regions. Generally in countries where the winter cold is intense, such as Canada, Scandinavia, and Finland, even the plains and lowlands are thickly strewn with innumerable ponds and small lakes, together with some others of a larger size; while in more temperate regions, such as Great Britain, Central and Southern Europe, the United States, and New Zealand, lake districts occur in all such mountainous tracts as can be proved to have been glaciated in times comparatively modern or since the geographical configuration of the surface bore a considerable resemblance to that now prevailing. In the same countries, beyond the glaciated regions, lakes abruptly cease, and in warmer and tropical countries are either entirely absent, or consist, as in equatorial Africa, of large sheets of water unaccompanied so far as we yet know by numerous smaller ponds and tarns.

The southern limits of the lake districts of the Northern Hemisphere are found at about 40° N. latitude on the American continent, and about 50° in Europe, or where the Alps intervene four degrees farther south. A large proportion of the smaller lakes are dammed up by barriers of unstratified drift, having the exact character of the moraines of glaciers, and are termed by geologists “morainic,” but some of them are true rock-basins, and would hold water even if all the loose drift now resting on their margins were removed.

In a paper read before the Geological Society of London in 1862, Professor Ramsay maintained that the first formation of most existing lakes took place during the glacial epoch, and was due, not to elevation or subsidence, but to actual erosion of their basins by glaciers. M. Mortillet in the same year advanced the theory that after the Alpine lake-basins had been filled up with loose fluviatile deposits, they were re-excavated by the great glaciers which passed down the valleys at the time of the greatest cold, a doctrine which would attribute to moving ice almost as great a capacity of erosion as that which assumed that the original basins were scooped out of solid rock by glaciers. It is impossible to deny that the mere geographical distribution of lakes points to the intimate connection of their origin with the abundance of ice during a former excess of cold, but how far the erosive action of moving ice has been the sole or even the principal cause of lake-basins, is a question still open to discussion.

The lakes of Switzerland and the north of Italy are some of them twenty and thirty miles in length, and so deep that their bottoms are in some cases from 1000 to 2000 feet beneath the level of the sea. It is admitted on all hands that they were once filled with ice, and as the existing glaciers polish and grind down, as before stated, the surface of the rocks, we are prepared to find that every lake-basin in countries once covered by ice should bear the marks of superficial glaciation, and also that the ice during its advance and retreat should have left behind it much transported matter as well as some evidence of its having enlarged the pre-existing cavity. But much more than this is demanded by the advocates of glacial erosion. They suggest that as the old extinct glaciers were several thousand feet thick, they were able in some places gradually to scoop out of the solid rock cavities twenty or thirty miles in length, and as in the case of Lago Maggiore from a thousand to two thousand six hundred feet below the previous level of the river-channel, and also that the ice had the power to remove from the cavity formed by its grinding action all the materials of the missing rocks. A constant supply, it is argued, of fine mud issues from the termination of every glacier in the stream which is produced by the melting of the ice, and this result of friction is exhibited both during winter and summer, affording evidence of the continual deepening and widening of the valleys through which glaciers pass. As the fine mud is carried away by a river from the deep pool which is formed from the base of every cataract, so it seems to be imagined that lake-basins may be gradually emptied of the mud formed by abrasion during the glacial period.

I am by no means disposed to object to this theory on the ground of the insufficiency of the time during which the extreme cold endured, but we must carefully consider whether that same time is not so vast as to make it probable that other forces, besides the motion of glaciers, must have cooperated in converting some parts of the ancient valley courses into lake-basins. They who have formed the most exalted conceptions of the erosive energy of moving ice do not deny that during the period termed “Glacial” there have been movements of the earth’s crust sufficient to produce oscillations of level in Europe amounting to 1000 feet or more in both directions. M. Charpentier, indeed, attributed some of the principal changes of climate in Switzerland, during the glacial period, to a depression of the central Alps to the extent of 3000 feet, and Swiss geologists have long been accustomed to attribute their lake basins, in part, to those convulsions by which the shape and course of the valleys may have been modified. Our experience, in the lifetime of the present generation, of the changes of level witnessed in New Zealand during great earthquakes is entirely opposed to the notion that the movements, whether upward or downward, are uniform in amount or direction throughout areas of indefinite extent. On the contrary, the land has been permanently raised in one region several feet or yards, and the rise has been found gradually to die out, so as to be imperceptible at a distance of twenty miles, and in some areas is even exchanged for a simultaneous downward movement of several feet.

But, it is asked, if such inequality of movement can have contributed towards the production of lake basins, does it not leave unexplained the comparative rarity of lakes in tropical and subtropical countries. In reply to this question it may be observed that in our endeavour to estimate the effects of subterranean movements in modifying the superficial geography of a country we must remember that each convulsion effects a very slight change. If it interferes with the drainage, whether by raising the lower or sinking the higher portion of a hydrographical basin, the upheaval or depression will only amount to a few feet at a time, and there may be an interval of years or centuries before any further movement takes place in the same region. In the mean time an incipient lake if produced may be filled up with sediment, and the recently-formed barrier will then be cut through by the river, whereas in a country where glacial conditions prevail no such obliteration of the temporary lake-basin would take place; for however deep it became by repeated sinking of the upper or rising of the lower extremity, being always filled with ice it might remain, throughout the greater part of its extent, free from sediment or drift until the ice melted at the close of the glacial period.

One of the most serious objections to the exclusive origin by ice-erosion of wide and deep lake-basins arises from their capricious distribution, as for example in Piedmont, both to the eastward and westward of Turin, where great lakes are wanting,[[6]] although some of the largest extinct glaciers descending from Mont Blanc and Monte Rosa came down from the Alps, leaving their gigantic moraines in the low country. Here, therefore, we might have expected to find lakes of the first magnitude rivalling the contiguous Lago Maggiore in importance.

A still more striking illustration of the same absence of lakes where large glaciers abound is afforded by the Caucasus, a chain more than 300 miles long, and the loftiest peaks of which attain heights from 16,000 to 18,000 feet. This greatest altitude is reached by Elbruz, a mountain in lat. 43° N. three degrees south of Mont Blanc, but on the other hand 3000 feet higher. The present Caucasian glaciers are equal or superior in dimensions to those of Switzerland, and like them give rise occasionally to temporary lakes by obstructing the course of rivers, and causing great floods when the icy barriers give way. Mr. Freshfield, a careful observer, writing in 1869, says:[[7]] “A total absence of lakes on both sides of the chains is the most marked feature. Not only are there no great subalpine sheets of water, like Como or Geneva, but mountain tarns, such as the Dauben See on the Gemmi, or the Klonthal See near Glarus, are equally wanting.” The same author states on the authority of the eminent Swiss geologist, Mons. E. Favre, who also explored the Caucasus in 1868, that moraines of great height and huge erratics of granite and other rocks “justify the assertion that the present glaciers of the Caucasus, like those of the Alps, are only the shadows of their former selves.”

It seems safe to assume that the chain of lakes, of which the Albert Nyanza forms one in equatorial Africa, was due to causes other than glacial. Yet if we could imagine a glacial period to visit that region filling the lakes with ice and scoring the rocks which form their sides and bottoms, we should be unable to decide how much the capacity of the basins had been enlarged and the surface modified by glacial erosion. The same may be true of the Lago Maggiore and Lake Superior, although the present basins of both of them afford abundant superficial markings due to ice-action.