NOTES.
Note 1 (p. 49).
I have been careful to speak of the "appearance" of cyclical evolution presented by living things; for, on critical examination, it will be found that the course of vegetable and of animal life is not exactly represented by, the figure of a cycle which returns into itself. What actually happens, in all but the lowest organisms, is that one part of the growing germ (A) gives rise to tissues and organs; while another part (B) remains in its primitive condition, or is but slightly modified. The moiety A becomes the body of the adult and, sooner or later, perishes, while portions of the moiety B are detached and, as offspring, continue the life of the species. Thus, if we trace back an organism along the direct line of descent from its remotest ancestor, B, as a whole, has never suffered death; portions of it, only, have been cast off and died in each individual offspring.
Everybody is familiar with the way in which the "suckers" of a strawberry plant behave. A thin cylinder of living tissue keeps on growing at its free end, until it attains a considerable length. At
successive intervals, it develops buds which grow into strawberry plants; and these become independent by the death of the parts of the sucker which connect them. The rest of the sucker, however, may go on living and growing indefinitely, and, circumstances remaining favourable, there is no obvious reason why it should ever die. The living substance B, in a manner, answers to the sucker. If we could restore the continuity which was once possessed by the portions of B, contained in all the individuals of a direct line of descent, they would form a sucker, or stolon, on which these individuals would be strung, and which would never have wholly died.
A species remains unchanged so long as the potentiality of development resident in B remains unaltered; so long, e.g., as the buds of the strawberry sucker tend to become typical strawberry plants. In the case of the progressive evolution of a species, the developmental potentiality of B becomes of a higher and higher order. In retrogressive evolution, the contrary would be the case. The phenomena of atavism seem to show that retrogressive evolution that is, the return of a species to one or other of its earlier forms, is a possibility to be reckoned with. The simplification of structure, which is so common in the parasitic members of a group, however, does not properly come under this head. The worm-like, limbless Lernoea has no resemblance to any of the stages of development of the many-limbed active animals of the group to which it belongs.