Bountifully to spread a table is much, but not enough, for without invitation how can hospitality be dispensed? To the feast of nectar the blossoms join their bidding; and those most conspicuously borne and massed, gayest of hue, richest in odor, secure most guests, and are therefore most likely to transmit to their kind their own excellences as hosts and entertainers. Thus all the glories of the blossoms have arisen in doing useful work; their beauty is not mere ornament, but the sign and token of duty well performed. Our opportunity to admire the radiancy and perfume of a jessamine or a pond-lily is due to the previous admiration of uncounted winged attendants. If a winsome maid adorns herself with a wreath from the garden, and carries a posy gathered at the brookside, it is for the second time that their charms are impressed into service; for the flowers' own ends of attraction all their scent and loveliness were called into being long before.
Let us put flowers of the blue flag beside those of the maple, and we shall have a fair contrast between the brilliancy of blossoms whose marrier has been an insect, and the dinginess of flowers indebted to the services of the wind. Can it be that both kinds of flowers are descended from forms resembling each other in want of grace and colour? Such, indeed, is the truth. But how, as the generations of the flowers succeeded one another, did differences so striking come about? In our rambles afield let us seek a clue to the mystery. It is late in springtime, and near the border of a bit of swamp we notice a clump of violets: they are pale of hue, and every stalk of them rises to an almost weedy height.
Wild Rose, Single
Twenty paces away, on a knoll of dry ground, we find more violets, but these are in much deeper tints of azure and yellow, while their stalks are scarcely more than half as tall as their brethren near the swamp. Six weeks pass by. This time we walk to a wood-lot close to a brimming pond. At its edge are more than a score wild-rose bushes. On the very first of them we see that some of the blossoms are a light pink, others a pink so deep as to seem dashed with vivid red. And while a flower here and there is decidedly larger and more vigorous than its fellows, a few of the blossoms are undersized and puny: the tide of life flows high and merrily in a fortunate rose or two, it seems to ebb and falter by the time it reaches one or two of their unhappy mates. As we search bush after bush we are at last repaid for sundry scratches from their thorns by securing a double rose, a “sport,” as the gardener would call it. And in the broad meadow between us and home we well know that for the quest we can have not only four-leaved clovers, but perchance a handful of five and six-leaved prizes. The secret is out. Flowers and leaves are not cast like bullets in rigid moulds, but differ from their parents much as children do. Usually the difference is slight, at times it is as marked as in our double rose. Whenever the change in a flower is for the worse, as in the sickly violets and roses we have observed, that particular change ends there—with death. But when the change makes a healthy flower a little more attractive to its insect ministers, it will naturally be chosen by them for service, and these choosings, kept up year after year, and century upon century, have at last accomplished much the same result as if the moth, the bee, and the rest of them had been given power to create blossoms of the most welcome forms, the most alluring tints, the most bewitching perfumes.
In farther jaunts afield we shall discover yet more. It is May, and a heavy rainstorm has caused the petals of a trillium to forget themselves and return to their primitive hue of leafy green. A month later we come upon a buttercup, one of whose sepals has grown out as a small but perfect leaf. Later still in summer we find a rose in the same surprising case, while not far off is a columbine bearing pollen on its spurs instead of its anthers. What family tie is betrayed in all this? No other than that sepals, petals, anthers and pistils are but leaves in disguise, and that we have detected nature returning to the form from which ages ago she began to transmute the parts of flowers in all their teeming diversity. The leaf is the parent not only of all these but of delicate tendrils, which save a vine the cost of building a stem stout enough to lift it to open air and sunshine. However thoroughly, or however long, a habit may be impressed upon a part of a plant, it may on occasion relapse into a habit older still, resume a shape all but forgotten, and thus tell a story of its past that otherwise might remain forever unsuspected. Thus it is with the somewhat rare “sport” that gives us a morning glory or a harebell in its primitive form of unjoined petals. The bell form of these and similar flowers has established itself by being much more effective than the original shape in dusting insect servitors with pollen. Not only the forms of flowers but their massing has been determined by insect preferences; a wide profusion of blossoms grow in spikes, umbels, racemes and other clusters, all economizing the time of winged allies, and attracting their attention from afar as scattered blossoms would fail to do. Besides this massing, we have union more intimate still as in the dandelion, the sun-flower and the marigold. These and their fellow composites each seem an individual; a penknife discloses each of them to be an aggregate of blossoms. So gainful has this kind of co-operation proved that composites are now dominant among plants in every quarter of the globe. As to how composites grew before they learned that union is strength, a hint is dropped in the “sport” of the daisy known as “the hen and chickens,” where perhaps as many as a dozen florets, each on a stalk of its own, ray out from a mother flower.
While for the most part insects have been mere choosers from among various styles of architecture set before them by plants, they have sometimes risen to the dignity of builders on their own account, and without ever knowing it. The buttress of the larkspur has sprung forth in response to the pressure of one bee's weight after another, and many a like structure has had the very same origin,—or shall we say, provocation? In these and in other examples unnumbered, culminating in the marvellous orchids and their ministers, there has come about the closest adaptation of flower-shape to insect-form, the one now clearly the counterpart of the other.
We must not forget that the hospitality of a flower is after all the hospitality of an inn-keeper who earns and requires payment. Vexed as flowers are apt to be by intruders that consume their stores without requital, no wonder that they present so ample an array of repulsion and defence. Best of all is such a resource as that of the red clover, which hides its honey at the bottom of a tube so deep that only a friendly bumblebee can sip it. Less effective, but well worth a moment's examination, are the methods by which leaves are opposed as fences for the discouragement of thieves. Here, in a Bellwort, is a perfoliate leaf that encircles the stem upon which it grows; and there in a Honeysuckle is a connate leaf on much the same plan, formed of two leaves, stiff and strong, soldered at their bases. Sometimes the pillager meets prickles that sting him, as in the roses and briers; and if he is a little fellow he is sure to regard him with intense disgust, a bristly guard of wiry hair—hence the commonness of that kind of fortification. Against enemies of larger growth a tree or shrub will often aim sharp thorns—another piece of masquerade, for thorns are but branches checked in growth, and frowning with a barb in token of disappointment at not being able to smile in a blossom. In every jot and tittle of barb and prickle, of the glossiness which disheartens or the gumminess which ensnares, we may be sure that equally with all the lures of hue, form and scent, nothing, however trifling it may seem, is as we find it, except through usefulness long tested and approved. In flowers, much that at first glance looks like idle decoration, on closer scrutiny reveals itself as service in disguise. In penetrating these disguises and many more of other phases, the student of flowers delights to busy himself. He loves, too, to detect the cousinship of plants through all the change of dress and habit due to their rearing under widely different skies and nurture, to their being surrounded by strangely contrasted foes and friends. Often he can link two plants together only by going into partnership with a student of the rocks, by turning back the records of the earth until he comes upon a flower long extinct, a plant which ages ago found the struggle for life too severe for it. He ever takes care to observe his flowers accurately and fully, but chiefly that he may rise from observation to explanation, from bare facts to their causes, from declaring What, to understanding, Whence and How.
One of the stock resources of novelists, now somewhat out of date, was the inn-keeper who beamed in welcome of his guest, grasped his hand in gladness, and loaded a table for him in tempting array, and all with intent that later in the day (or night) he might the more securely plunge a dagger into his victim's heart—if, indeed, he had not already improved an opportunity to offer to that victim's lips a poisoned cup. This imagined treachery might well have been suggested by the behaviour of certain alluring plants that so far from repelling thieves, or discouraging pillagers, open their arms to all comers—with purpose of the deadliest. Of these betrayers the chief is the round-leaved sun-dew, which plies its nefarious vocation all the way from Labrador to Florida. Its favourite site is a peat-bog or a bit of swampy lowland, where in July and August we can see its pretty little white blossoms beckoning to wayfaring flies and moths their token of good cheer! Circling the flower-stalk, in rosette fashion, are a dozen or more round leaves, each of them wearing scores of glands, very like little pins, a drop of gum glistening on each and every pin by way of head. This appetizing gum is no other than a fatal stick-fast, the raying pins closing in its aid the more certainly to secure a hapless prisoner. Soon his prison-house becomes a stomach for his absorption. Its duty of digestion done, the leaf in all seeming guilessness once more expands itself for the enticement of a dupe. To see how much the sun-dew must depend upon its meal of insects we have only to pull it up from the ground. A touch suffices—it has just root enough to drink by; the soil in which it makes, and perhaps has been obliged to make, its home has nothing else but drink to give it.
Less accomplished in its task of assassination is the common butterwort to be found on wet rocks in scattered districts of Canada and the States adjoining Canada. Surrounding its pretty violet flowers, of funnel shape, are gummy leaves which close upon their all too trusting guests, but with less expertness than the sun-dew's. The butterwort is but a 'prentice hand in the art of murder, and its intended victims often manage to get away from it. Built on a very different model is the bladderwort, busy in stagnant ponds near the sea coast from Nova Scotia to Texas. Its little white spongy bladders, about a tenth of an inch across, encircle the flowering stem by scores. From each bladder a bunch of twelve or fifteen hairy prongs protrude, giving the structure no slight resemblance to an insect form. These prongs hide a valve which, as many an unhappy little swimmer can attest, opens inward easily enough, but opens outward never. As in the case of its cousinry a-land, the bladderwort at its leisure dines upon its prey.