The evolution of organic matter now claims attention in detail. Of the origin of first life, we know absolutely nothing. The doctrine of Evolution does not deal with that. There are, however, many hypotheses upon the subject. Lord Kelvin, the eminent physicist, has suggested that unicellular life may have been transferred to this globe from a wrecked planet. This hypothesis obviously aids us very little, for it merely transfers the original scene of action to some other world. Personally, I prefer the idea that the first protoplasm was produced by the action of the sun upon inorganic matter not unlike the colloids, and that it “fed upon the previous steps in its own evolution.” In this connection, I may say that two points are certain—viz., that vegetable life preceded animal life, and that the first forms of life were mere specks of jelly, without organs. Can these primitive specks be created at the present time? Or, in other words, can protoplasm be manufactured by artificial processes? The answer must be No; not by any process now known, although a great number of experiments have been made with the object of manufacturing unicellular vegetable life. During the years between 1870 and 1880, this question was thoroughly thrashed out, and at first the balance seemed to be very evenly held between the supporters and the opponents of spontaneous generation. The investigations of the late Professor Tyndall, however, conclusively proved that biogenesis, that is, all life from previous life, is the condition at the present day. But I must add Huxley’s words of warning, viz., “that with organic chemistry, molecular physics, and physiology yet in their infancy, and every day making prodigious strides, it would be the height of presumption for any man to say that the conditions under which matter assumes the qualities called vital, may not some day be artificially brought together.” And further, “that as a matter not of proof but of probability, if it were given me to look beyond the abyss of geologically recorded time, to the still more remote period when the earth was passing through chemical and physical conditions which it can never see again, I should expect to be a witness of the evolution of living protoplasm from nonliving matter.”

The first protoplasm must be extremely ancient, for the remains of sea-weeds are found in the oldest strata, and vegetation implies the manufacture of protoplasm from inorganic matter.

When the earth was in the condition to which Huxley referred, the constantly decreasing heat, and the recurrence of the seasons produced, by slow degrees, changes in the congenital character of the forms of life. Every individual varied somewhat from its predecessors, and those forms which possessed variations most suitable to the environment were the ones which eventually survived. The transition from the protophyta, the lowest class of vegetable life, to the protozoa, the lowest class of animal life, must have been a very simple matter in the condition in which the earth then was. Indeed, today the difference between the lowest microscopic animals and the lowest microscopic plants is by no means clearly defined.

Innumerable hosts of life made their appearance upon our planet while the surface was going through the cooling process, and they were, at first, of course, of the most primitive kind. But the same laws were always at work, viz., no two living things were exactly alike when they made their appearance upon this earth, although the differences between several forms might be very slight. Variation was, and is, the order of the day.

The individuals which possessed variations in accordance with the environment persisted, while those having injurious variations had a tendency to disappear. Congenital variations were (and are) transmitted with great certainty. This is Mr. Darwin’s “Process of Natural Selection,” called by Mr. Spencer “The Survival of the Fittest.”

The other Darwinian factor in evolution is Sexual Selection. It is that department of Natural Selection in which sex is especially concerned. Anything which exhibits the prowess or beauty of the one sex attracts the other, and decides the preference for one individual over another, with the result that those individuals which are unattractive to the opposite sex are unable to reproduce their kind. The importance of this factor will be appreciated if I give an extract from Darwin’s “Descent of Man” (Vol. II., p. 367). “For my own part,” wrote our great master, “I conclude that of all the causes which have led to the differences in external appearance between the races of men, and to a great extent between man and the lower animals, sexual selection must have been by far the most efficient.”

As I have already said, Darwin neither invented nor discovered the doctrine of Evolution. But he placed it upon a firm foundation by the discovery of the two great factors to which I have referred, and, by incessant observation and indomitable energy, he demonstrated the truth of them beyond any reasonable doubt.

The proofs of the truth of Evolution are of two kinds—palaeontological and embryological. The palaeontological evidence has found its way into popular books, and even into some of the literary newspapers. The history of the horses, of the crocodiles, of the rhinoceros is known in detail. All the stages have been found which intervene between the four-toed Eohippos of the Lower Eocene and the zebra and horse of the present day. Thanks to the late Professor Marsh, of Yale, not only are the successive steps in the evolution of the foot-structure preserved, but so also are the various stages in the evolution of the teeth. The occasional appearance of a three-toed horse points very plainly to a three-toed progenitor, a striking example of atavism, that is, the reappearance of a characteristic which has “skipped” one or more generations.

If the principle of heredity be true, one would expect to find in the development of animals and plants, traces of the line of descent. “If Evolution be true, one ought to find, following back the development of the egg, that specific details would vanish and give rise to more generalized features; that the earlier the stages, the more the embryos of related forms would resemble each other.” This is exactly what is found, there being, in a vast number of instances, a remarkable parallel between the palaeontological record and the embryological evidence. A detailed examination of the facts would not be intelligible to anybody who is not a practical biologist; but I am fully warranted in asserting that every organism in the course of its life-history (technically called ontogeny) is a recapitulation of the history of the race—technically known as phylogeny.

There is other evidence in abundance. The phenomena named atavism is a part of that evidence. Almost everybody has seen well-defined and regular stripes upon horses, and nobody doubts that they indicate a zebra-like ancestor. Again, in the inner side of the human eye is a little red fold, known as the plica semilunaris, the remnant of an ancestor which possessed a third eyelid, similar to that possessed by some reptiles and birds of to-day.