Who are the supporters of the doctrine of Evolution? Practically the whole scientific world. The late Professor Marsh, the distinguished palaeontologist, when president of the American Association for the Advance of Science in 1878, said:

“I need offer no argument for Evolution, since to doubt evolution is to doubt science, and science is only another name for the truth.” Professor Marsh meant, of course, not that evolution is to be taken “on trust,” but that it has been so thoroughly proved that new arguments in support of it are unnecessary.

Concerning Natural Selection, sometimes called Darwinism, the late Professor Huxley said (quotation from Darwin’s “Life”): “I venture to affirm that so far as all my knowledge goes, all the ingenuity and all the learning of the hostile critics have not enabled them to adduce a single fact of which it can be said this is irreconcilable with the Darwinian theory.”

I occasionally hear the old argument that species are immutable—that a species is something which never changes. It seems a little late in the day to revive this contention, but it is necessary to be prepared with a reply. The critics of Darwin’s theory of “the Origin of Species by Natural Selection” have always refused to give a tangible definition of the word “species,” and, as a result, the real difficulty turns upon that point. What is a species? Linnaeus said: “There are as many species as an infinite Being created at the beginning,” a statement which is a confession of faith, and not a scientific definition. We must remember, of course, that Linnaeus died as long ago as 1778. The truth is that all the various tests for species have proved faulty, that of the fertility of hybrids having little more value than many of the other so-called “tests.” In classification, the word “species” means the lowest subdivision to which a name is usually applied, and to aid the zoologist’s or botanist’s memory, some system of classification is, I need not say, an absolute necessity.

According to the view of the anti-evolutionists, most of whom are not scientific men, descendants of a common ancestor must belong to the same species. Nevertheless, the late Mr. Romanes has shown that the rabbits of Porto Santo, an island in the Atlantic, about twenty-five miles from Madeira, descended from the European stock of nearly 500 years ago, will no longer breed with their continental cousins.

When we remember that some wild animals will not breed in captivity, the idea of sterility as a test of species seems utterly unscientific. I venture to say that there can be no accurate definition of species in terms of physiology, for every individual has its peculiarities, chemical as well as physical, and the real difficulty is to decide when these peculiarities are important enough to make it useful to give a precise name to their possessors. Assume for a moment that a species is a group of individuals agreeing in essential characters which remain constant from one generation to another. But what are essential characters and how much constancy is demonstrated? Upon these points no two biologists are likely to agree. For example, taking the birds of Germany, Bechstein says there are 367 species; Brehm says there are 900. According to Reichenbach there are 379, and Meyer and Wolf tell us there are 406.

The idea of a species is based upon structural resemblances between individuals, and the degree of importance attached to these depends upon the mind of the particular observer.

There are two reasons why nobody has seen one species turn into another. The first is that until the word “species” is satisfactorily defined, instances of the evolution of new forms cannot be supplied. Secondly, as nobody lives much beyond a hundred years at the most—a mere moment in Nature—our ability to witness marked changes in animals or plants is extremely limited. Minor changes, of course, are frequently noticed. I ask the reader to remember, however, that the flower-garden and the farm-yard are in an artificial condition, Natural Selection having ceased. For instance, the duck which has defective wings when hatched has as good a chance of surviving as the duck with powerful wings.

Who are the opponents of the doctrine of Evolution? In the scientific world they are difficult to find. Professor Virchow, of Berlin, the distinguished pathologist must, I think, be classed as one, although his verdict is really “not proven.” Professor Haeckel, however, has pointed out that the opinion of a pathologist, no matter how eminent, upon the subject of evolution cannot carry much weight.

Until recently we had with us two men of science whose opposition to some portion of the doctrine of evolution was of importance. These men were Sir William Dawson, the Canadian geologist, and Mr. Mivart, the English anatomist. Both of these gentlemen have died within the past two years.