The genus Rhinosporidium should probably be placed in a distinct family. The only species so far described is R. kinealyi from the nasal septum of man, to which reference has above been made. Another form, Neurosporidium cephalodisci, agreeing in some respects with Rhinosporidium, has been described by Ridewood and Fantham (37a) from the nervous system of Cephalodiscus.

A parasite whose affinities are doubtful, but which is regarded by Caullery and Mesnil as allied to the Haplosporidia, is the curious parasite originally described by Schewiakoff as “endoparasitic tubes” of Cyclops; it has been named by Caullery and Mesnil, Scheviakovella. This organism is remarkable in one or two ways: it possesses a contractile vacuole; the amoeboid trophozoites tend to form plasmodia; and the spores, of the usual simple type, may apparently divide by binary fission.

5. There remain, lastly, certain forms, which are conveniently grouped together as “Sporozoa incertae sedis,” either for the reason that it is impossible to place them in any of the well-defined orders, or because their life-cycle is at present too insufficiently known. Serosporidia is the name given by Pfeiffer to certain minute parasites of the body-cavity of Crustacea; they include Serosporidium, Blanchardina and Botellus. Lymphosporidium, a form with distributed nucleus, causing virulent epidemics among brook-trout, is considered by Calkins(3) to be suitably placed here. Another parasite of lymphatic spaces and channels is the remarkable Lymphocystis, described by Woodcock (46), from plaice and flounders, which in some respects rather recalls a Gregarine. The group Exosporidia was founded by Perrier to include a peculiar organism, ectoparasitic on Arthropods, to which the name of Amoebidium had been given by Cienkowsky. It has recently been shown, however, that this organism is most probably an Alga. Another genus, Exosporidium, described by Sand (38), is placed at present in this group. For details of the structure of these forms and others like Siedleckia, Toxosporidium, Chitonicium Joyeuxella and Metschnikovella, a comprehensive treatise on the Sporozoa, such as that of Minchin, should be consulted.

To complete this article, it will be sufficient to mention various enigmatical bodies, associated with different diseases, which are regarded by their describers as Protozoa. Among such is the “Histosporidium carcinomatosum” of Feinberg, which he finds in cancerous growths. Cytoryctes, the name given to “Guarnieri’s bodies” in small-pox and vaccinia, has been recently investigated by Calkins (3a), who has described a complex life-cycle for the alleged parasite. Other workers, however, such as Siegel, give a quite different account of these bodies, and, moreover, find similar ones in scarlet-fever, syphilis, &c.; while yet others (e.g. Prowazek) deny that they are parasitic organisms at all.

Bibliography.—(For general works see under [Sporozoa].) (1) Bertram, “Beiträge zur Kenntnis der Sarcosporidien,” Zool. Jahrb. Anat. 5, 1902; (2) L. Brasil, “Joyeuxella toxoides,” (n.g., n.sp.), Arch. zool. exp. N. et R. (3) 10, p. 5, 7 figs., 1902; (3) G.N. Calkins, “Lymphosporidium truttae,” (n.g., n.sp.), Zool. Anz. 23, p. 513, 6 figs., 1903; (3a) ib. The Life-History of Cytoryctes Variolae; Guarnieri, “Studies path. etiol. variola,” J. Med. Research (Boston, 1904), p. 136, 4 pls.; (3b) M. Caullery and A. Chappellier, “Anurosporidium pelseneeri, (n.g., n.sp.), Haplosporidie,” &c., C. R. soc. biol. 60, p. 325, 1906; (4) M. Caullery and F. Mesnil, “Sur un type nouveau” (Metchnikovella, n.g.), C. R. ac. sci. 125, p. 787, 10 figs., 1897; (5) ib. “Sur trois Sporozoaires parasites de la Capitella,” C. R. soc. biol. 49, p. 1005, 1877; (6) ib. “Sur un Sporozoaire aberrant” (Siedleckia, n.g.), op. cit. 50, p. 1093, 7 figs., 1898; (7) ib. “Sur le genre Aplosporidium” (nov.), op. cit. 51, p. 789, 1899; (8) ib. “Sur les Aplosporidies,” C. R. ac. sci. 129, p. 616, 1899; (9) ib. “Sur les parasites intimes des Annélides” (Siedleckia, Toxosporidium), C. R. ass. franç., 1899, p. 491, 1900; (10) ib. “Sur un type nouveau (Sphaeractinomyxon, n.g.) d’Actinomyxidies,” C. R. soc. biol. 56, p. 408, 1904; (11) ib. “Phénomènes de sexualité dans le développement des Actinomyxidies,” op. cit. 58, p. 889, 1905; (12) ib. “Recherches sur les Actinomyxidies,” Arch. Protistenk. 6, p. 272, pl. 15, 1905; (13) ib. “Sur quelques nouvelles Haplosporidies d’Annélides,” C. R. soc. biol. 58, p. 580, 6 figs., 1905; (14) ib. “Sur des Haplosporidies parasites de poissons marins,” ib. p. 640, 1905; (15) ib. “Recherches sur les Haplosporidies,” Arch. zool. exp. (4) 4, p. 101, pls. 11-13, 1905; (16) L. Cohn, “Über die Myxosporidien von Esox lucius,” Zool. Jahr. Anat. 9, p. 227, 2 pls., 1896; (17) ib. “Zur Kenntniss der Myxosporidien,” Centrbl. Bakt. 1, Orig. 32, p. 628, 3 figs., 1902; (18) ib. “Protozoen als Parasiten in Rotatorien,” Zool. Anz. 25, p. 497, 1902; (19) F. Doflein, “Über Myxosporidien,” Zool. Jahr. Anat. 11, p. 281, 6 pls., 1898; (20) ib. “Fortschritte auf dem Gebiete der Myxosporidienkunde,” Zool. Centrbl. 7, p. 361, 1899; (21) R. Gurley, “The Myxosporidia,” Bull. U.S. Fish. Comm., 1892, p. 65, 47 pls., 1894; (22) E. Hesse, “Sur une nouvelle Microsporidie tétrasporée du genre Gurleya,” C. R. soc. biol. 55, p. 495, 1903; (23) ib. “Thelohania légeri” (n.sp.), op. cit. 57, pp. 570-572, 10 figs., 1904; (24) ib. “Sur Myxocystis Mrazeki Hesse,” &c., op. cit. 58, p. 12, 9 figs., 1905; (25) A. Laveran and F. Mesnil, “Sur la multiplication endogène des Myxosporidies,” op. cit. 54, p. 469, 5 figs., 1902; (26) ib. “Sur la morphologie des Sarcosporidies,” op. cit. 51, p. 245, 1899; (27) ib. “De la Sarcocystin,” op. cit. p. 311, 1899; (28) L. Léger, “Sur la sporulation du Triactinomyxon,” op. cit. 56, p. 844, 4 figs., 1904; (29) ib. “Considérations sur ... les Actinomyxidies,” op. cit. p. 846, 1904; (29a) L. Léger and E. Hesse, “Sur une nouvelle Myxosporidie, Coccomyxa, n.g.,” C. R. ac. sci., 1st July 1907; (29b) ib. “Sur la structure de la paroisporale des Myxosporidies,” op. cit. 142, p. 720, 1906; (29c) A. Lutz and A. Splendore, “Über ‘Pébrine’ and verwandte Mikrosporidien,” Centrbl. Bakt. 1, 33, Orig. p. 150, 1903, and 36, Orig. p. 645, 2 pls., 1904; (29d) E.A. Minchin and H.B. Fantham, “Rhinosporidium kinealyi” (n.g., n.sp.), Q. J. Micr. Sci. 49, p. 521, 2 pls., 1905; (30) A. Mrazek, “Über eine neue Sporozoenform” (Myxocystis), S. B. Böhm. Ges. 8, 5 pp., 9 figs., 1897; (31) ib. “Glugea lophii,” Doflein, op. cit. 10, 8 pp., 1 pl., 1899; (32) C. Perez, “Sur un organisme nouveau, Blastulidium,” C. R. soc. biol. 55, p. 715, 5 figs., 1903; (33) ib. “Sur nouvelles Glugéidées,” op. cit. 58, pp. 146-151, 1905; (34) ib. “Microsporidies parasites des crabes,” Bull. sta. biol. d’Arcachon, 8, 22 pp., 14 figs., 1905; (35) W.S. Perrin, “Pleistophora periplanetae,” Q. J. Micr. Sci. 49, p. 615, 2 pls., 1906; (36) L. Plate, “Über einen einzelligen Zellparasiten” (Chitonicium), Fauna Chilensis, 2, pp. 601, pls., 1901; (37) M. Plehn, “Über die Drehkrankheit der Salmoniden” (Lentospora, n.g.), Arch. Protistenk. 5, p. 145, pl. 5, 1904; (37a) W.J. Ridewood and H.B. Fantham, “Neurosporidium cephalodisci, n.g., n.sp.,” Q. J. Micr. Sci. 51, p. 81, pl. 7, 1907; (38) R. Sand, “Exosporidium marinum” (n.g., n.sp.), Bull. soc. micr. belge, 24, p. 116, 1898; (39) T. Smith, “The production of sarcosporidiosis in the mouse,” &c., J. Exp. Med. 6, p. 1, 4 pls., 1901; (40) W. Stempell, “Über Thelohania mülleri,” Zool. Jahr. Anat. 16, p. 235, pl. 25, 1902; (41) ib. “Über Polycaryum branchiopodianum” (n.g., n.sp.), Zool. Jahrb. Syst. 15, p. 591, pl. 31, 1902; (42) ib. “Über Nosema anomalum,” Arch. Protistenk, 4, p. 1, pls. 1-3, 1904; (43) P. Thélohan, “Recherches sur les Myxosporidies,” Bull. sci. France belg. 26, p. 100, 3 pls., 1895; (44) P. Vuillemin, “Le Sarcocystis tenella, parasite de l’homme,” C. R. ac. sci. 134, p. 1152, 1902; (45) H.M. Woodcock, “On Myxosporidia in flat fish,” Proc. Liverp. Biol. Soc. 18, p. 126, pl. 2, 1904; (46) ib. “On a remarkable parasite” (Lymphocystis), op. cit. p. 143, pl. 3, 1904.

(H. M. Wo.)


ENDYMION, in Greek mythology, son of Aëthlius and king of Elis. He was loved by Selene, goddess of the moon, by whom he had fifty daughters, supposed to represent the fifty moons of the Olympian festal cycle. In other versions, Endymion was a beautiful youth, a shepherd or hunter whom Selene visited every night while he lay asleep in a cave on Mount Latmus in Caria (Pausanias v. 1; Ovid, Ars am. iii. 83). Zeus left him free to choose anything he might desire, and he chose an everlasting sleep, in which he might remain youthful for ever (Apollodorus i. 7). According to others, Endymion’s eternal sleep was a punishment inflicted by Zeus upon him because he ventured to fall in love with Hera, when he was admitted to the society of the Olympian gods (Schol. Theocritus iii. 49). The usual form of the legend, however, represents Endymion as having been put to sleep by Selene herself in order that she might enjoy his society undisturbed (Cicero, Tusc. disp. i. 38). Some see in Endymion the sun, setting opposite to the rising moon, the Latmian cave being the cave of forgetfulness, into which the sun plunges beneath the sea; others regard him as the personification of sleep or death (see Mayor on Juvenal x. 318).


ENERGETICS. The most fundamental result attained by the progress of physical science in the 19th century was the definite enunciation and development of the doctrine of energy, which is now paramount both in mechanics and in thermodynamics. For a discussion of the elementary ideas underlying this conception see the separate heading [Energy].