Characteristics of the root.—The roots of the tomato plant, while abundant in number, are short and can only gather food and water from a limited area. A plant of garden bean, for instance, is not more than half the size of one of the tomato, but its roots extend through the soil to a greater distance, gather plant food from a greater bulk of soil, seem better able to search out and gather the particular food element which the plant needs than do those of the tomato. This characteristic of the latter plant makes the composition of the soil as to the proportion of easily available food elements of great importance. Tomato roots are also exceedingly tender and incapable of penetrating a hard and compact soil, so that the condition of the soil as to tilth is of greater importance with regard to tomatoes than with most garden vegetables.
Another characteristic of the tomato roots is that the period of their active life is short. When young they are capable of transmitting water and nutritive material very rapidly, but they soon become clogged and inefficient to such an extent as to result in the starvation and death of the plant. If the branches of such an exhausted plant be bent over and covered with earth they will frequently start new roots and produce a fresh crop of fruit, or if plants which have made a crop in the greenhouse be transplanted to the garden and cut back, a new set of roots will often develop and the plant will produce a second crop of fruit which, in amount, often equals or exceeds the first one. But such growths come only from new roots springing from the stem—never from an extension of the old root system.
Characteristics of the stem and leaves.—The growth of the stem, and leaves of the young tomato plant is very rapid and, the cellular structure coarse, loose and open. A young branch is easily broken and when this is done it shows scarcely any fibrous structure—simply a mass of coarse cellular matter which while capable, when young, of transmitting nutritive matter rapidly, soon becomes dogged and inert. This structure not only makes the active life of the leaves short, like that of the roots, but necessitates a fresh growth in order to continue the fruitfulness of the plant and renders the leaves very susceptible to injury from bacterial and fungous diseases. The rapid growth also necessitates an abundance of sunlight.
Characteristics of the blossom.—The inflorescence of the tomato is usually abundant and it is rare that a plant does not produce sufficient blooms for a full crop. The flowers are perfect as far as parts are concerned (Fig. 2) and in bright, sunny weather there is an abundance of pollen, but sunlight and warmth are essential to its maturing into a condition in which it can easily reach the stigma. The structure and development of the flower are such that while occasionally, particularly in healthy plants out of doors, the stigma becomes receptive and takes the pollen as it is pushed out through the stamen tube by the elongating style, it is more often pushed beyond them before the pollen matures, so that the pollen has to reach the stigma through some other means. Usually this is accomplished by the wind, either directly or through the motion of the plants.
Under glass it is generally necessary to assist the fertilization either directly by application or by motion of the plant, this latter only being effective in the middle of a bright sunny day. In the open ground in cold, damp weather the flowers often fail of fertilization, in which case they drop, and this is often the first indication of a failing of the crop on large, strong vines. I have known of many cases where the yield of fruit from large and seemingly very healthy vines was very light because continual rains prevented the pollenization of the flowers. Such failures, however, do not always come from a want of pollen but may result from an over or irregular supply of water either at the root or in the air, imperfectly balanced food supply, a sapping of the vitality of the plants when young, or from other causes. Insects rarely visit tomato flowers and are seldom the means of their fertilization.
Characteristics of the fruit.—The fruit of the original species from which our cultivated tomatoes have developed was doubtless a comparatively small two to many-celled berry, with comparatively dry central placenta and thin walls. In some species the cells were indicated by distinct sutures, forming a rough or corrugated fruit. It has improved under cultivation by increase in size, the material thickening of the cell walls, the development of greater juiciness and richer flavor and a decrease in the size and dryness of the placenta, as well as the breaking up of the cells by fleshy partitions resulting in the disappearance of the deep sutures and an improvement in the smoothness and beauty of the fruit. (Fig. 11.)
The quality of the fruit is largely dependent upon varietal differences, to be spoken of later, but it is also influenced by conditions of growth—such as the proportion of the nutritive elements found in the soil, the proper supply of moisture, the degree and uniformity of temperature and, most of all, the amount of sunlight. Sudden changes of temperature and moisture often result in cracks and fissures in the skin and flesh, which not only injure the appearance but affect the flavor of the fruit.
FIG. 11—TYPICAL BUNCH OF MODERN TOMATOES
Contrast with Figs. 9 and 10