Fig. 2.
Thus, a tired wayfarer on a hot day throws himself on the damp earth beneath a maple-tree. The sensations of delicious rest and coolness pouring themselves through the direct line would naturally discharge into the muscles of complete extension: he would abandon himself to the dangerous repose. But the loop-line being open, part of the current is drafted along it, and awakens rheumatic or catarrhal reminiscences, which prevail over the instigations of sense, and make the man arise and pursue his way to where he may enjoy his rest more safely. Presently we shall examine the manner in which the hemispheric loop-line may be supposed to serve as a reservoir for such reminiscences as these. Meanwhile I will ask the reader to notice some corollaries of its being such a reservoir.
First, no animal without it can deliberate, pause, postpone, nicely weigh one motive against another, or compare. Prudence, in a word, is for such a creature an impossible virtue. Accordingly we see that nature removes those functions in the exercise of which prudence is a virtue from the lower centres and hands them over to the cerebrum. Wherever a creature has to deal with complex features of the environment, prudence is a virtue. The higher animals have so to deal; and the more complex the features, the higher we call the animals. The fewer of his acts, then, can such an animal perform without the help of the organs in question. In the frog many acts devolve wholly on the lower centres; in the bird fewer; in the rodent fewer still; in the dog very few indeed; and in apes and men hardly any at all.
The advantages of this are obvious. Take the prehension of food as an example and suppose it to be a reflex performance of the lower centres. The animal will be condemned fatally and irresistibly to snap at it whenever presented, no matter what the circumstances may be; he can no more disobey this prompting than water can refuse to boil when a fire is kindled under the pot. His life will again and again pay the forfeit of his gluttony. Exposure to retaliation, to other enemies, to traps, to poisons, to the dangers of repletion, must be regular parts of his existence. His lack of all thought by which to weigh the danger against the attractiveness of the bait, and of all volition to remain hungry a little while longer, is the direct measure of his lowness in the mental scale. And those fishes which, like our cunners and sculpins, are no sooner thrown back from the hook into the water, than they automatically seize the hook again, would soon expiate the degradation of their intelligence by the extinction of their type, did not their exaggerated fecundity atone for their imprudence. Appetite and the acts it prompts have consequently become in all higher vertebrates functions of the cerebrum. They disappear when the physiologist's knife has left the subordinate centres alone in place. The brainless pigeon will starve though left on a corn-heap.
Take again the sexual function. In birds this devolves exclusively upon the hemispheres. When these are shorn away the pigeon pays no attention to the billings and cooings of its mate. And Goltz found that a bitch in heat would excite no emotion in male dogs who had suffered large loss of cerebral tissue. Those who have read Darwin's 'Descent of Man' know what immense importance in the amelioration of the breed in birds this author ascribes to the mere fact of sexual selection. The sexual act is not performed until every condition of circumstance and sentiment is fulfilled, until time, place, and partner all are fit. But in frogs and toads this passion devolves on the lower centres. They show consequently a machine-like obedience to the present incitement of sense, and an almost total exclusion of the power of choice. Copulation occurs per fas aut nefas, occasionally between males, often with dead females, in puddles exposed on the highway, and the male may be cut in two without letting go his hold. Every spring an immense sacrifice of batrachian life takes place from these causes alone.
No one need be told how dependent all human social elevation is upon the prevalence of chastity. Hardly any factor measures more than this the difference between civilisation and barbarism. Physiologically interpreted, chastity means nothing more than the fact that present solicitations of sense are overpowered by suggestions of æsthetic and moral fitness which the circumstances awaken in the cerebrum; and that upon the inhibitory or permissive influence of these alone action directly depends.
Within the psychic life due to the cerebrum itself the same general distinction obtains, between considerations of the more immediate and considerations of the more remote. In all ages the man whose determinations are swayed by reference to the most distant ends has been held to possess the highest intelligence. The tramp who lives from hour to hour; the bohemian whose engagements are from day to day; the bachelor who builds but for a single life; the father who acts for another generation; the patriot who thinks of a whole community and many generations; and finally, the philosopher and saint whose cares are for humanity and for eternity,—these range themselves in an unbroken hierarchy, wherein each successive grade results from an increased manifestation of the special form of action by which the cerebral centres are distinguished from all below them.
In the 'loop-line' along which the memories and ideas of the distant are supposed to lie, the action, so far as it is a physical process, must be interpreted after the type of the action in the lower centres. If regarded here as a reflex process, it must be reflex there as well. The current in both places runs out into the muscles only after it has first run in; but whilst the path by which it runs out is determined in the lower centres by reflections few and fixed amongst the cell-arrangements, in the hemispheres the reflections are many and instable. This, it will be seen, is only a difference of degree and not of kind, and does not change the reflex type. The conception of all action as conforming to this type is the fundamental conception of modern nerve-physiology. So much for our general preliminary conception of the nerve-centres! Let us define it more distinctly before we see how well physiological observation will bear it out in detail.