At the time when Darwin's work appeared there already existed, as we have seen, a fully formed morphology with set and definite principles. The aim of this pre-evolutionary morphology had been to discover and work out in detail the unity of plan underlying the diversity of forms, to disentangle the constant in animal form and distinguish from it the accessory and adaptive. The main principle upon which this work was based was the principle of connections, so clearly stated by Geoffroy. The principle of connections served as a guide in the search for the archetype, and this search was prosecuted in two directions—first, by the comparison of adult structure; and second, by the comparative study of developing embryos. It was found that the archetype was shown most clearly by the early embryo, and this embryological archetype came to be preferred before the archetype of comparative anatomy. It became apparent also that the parts first formed (germ-layers) were of primary importance for the establishing of homologies.
While practically all morphologists were agreed as to the main principles of their science, they yet showed, as regards their general attitude to the problems of form, a fairly definite division into two groups, of which one laid stress upon the intimate relation existing between form and function, while the other disregarded function completely, and sought to build up a "pure" or abstract morphology. In opposition to both groups, in opposition really to morphology altogether, a movement had gained strength which tended towards the analysis and disintegration of the organism. This movement took its origin in the current materialism of the day, and found expression particularly in the cell-theory and in materialistic physiology.
The separation between morphology as the science of form and physiology as the science of the physics and chemistry of the living body had by Darwin's day become well-nigh absolute.
The morphology of the 'fifties lent itself readily to evolutionary interpretation. Darwin found it easy to give a formal solution of all the main problems which pre-evolutionary morphology had set—he was able to interpret the natural system of classification as being in reality genealogical, systematic relationship as being really blood-relationship; he was able to interpret homology and analogy in terms of heredity and adaptation; he was able to explain the unity of plan by descent from a common ancestor, and for the concept of "archetype" to substitute that of "ancestral form."
The current morphology, Darwin found, could be taken over, lock, stock and barrel, to the evolutionary camp.
In what follows we shall see that the coming of evolution made surprisingly little difference to morphology, that the same methods were consciously or unconsciously followed, the same mental attitudes taken up, after as before the publication of the Origin of Species.
Darwin himself was not a professional morphologist; the conversion of morphology to evolutionary ideas was carried out principally by his followers, Ernst Haeckel and Carl Gegenbaur in Germany, Huxley, Lankester, and F. M. Balfour in England.
It was in 1866 that Haeckel's chief work appeared, a General Morphology of Organisms,[366] which was intended by its author to bring all morphology under the sway and domination of evolution.
It was a curious production, this first book of Haeckel's, and representative not so much of Darwinian as of pre-Darwinian thought. It was a medley of dogmatic materialism, idealistic morphology, and evolution theory; its sources were, approximately, Büchner, Theodor Schwann, Virchow, H. G. Bronn, and, of course, Charles Darwin.
It was scarcely modern even on its first appearance, and many regarded it, not without reason, as a belated offshoot of Naturphilosophie.