The Four Great Linkage Groups of Drosophila Ampelophila
The following description of the characters of the wild fly may be useful in connection with the account of the modifications of these characters that appear in the mutants.
The head and thorax of the wild fly are grayish-yellow, the abdomen is banded with alternate stripes of yellow and black. In the male, (fig. 4 to right), there are three narrow bands and a black tip. In the female there are five black bands (fig. 4 to left). The wings are gray with a surface texture of such a kind that at certain angles they are iridescent. The eyes are a deep, solid, brick-red. The minute hairs that cover the body have a very definite arrangement that is most obvious on the head and thorax. There is a definite number of larger hairs called bristles or chaetae which have a characteristic position and are used for diagnostic purposes in classifying the species. On the foreleg of the male there is a comb-like organ formed by a row of bristles; it is absent in the female. The comb is a secondary sexual character, and it is, so far as known, functionless.
Some of the characters of the mutant types are shown in figures 53, 54, 55, 56. The drawing of a single fly is often used here to illustrate more than one character. This is done to economize space, but of course there would be no difficulty in actually bringing together in the same individual any two or more characters belonging to the same group (or to different groups). Without colored figures it is not possible to show many of the most striking differences of these mutant races; at most dark and light coloring can be indicated by the shading of the body, wings, or eyes.
Group I
In the six flies drawn in figure 53 there are shown five different wing characters. The first of these types (a) is called cut, because the ends of the wings look as though they had been cut to a point. The antennae are displaced downward and appressed and their bristle-like aristae are crumpled.
The second figure (b) represents a fly with a notch in the ends of the wings. This character is dominant, but the same factor that
produces the notch in the wings is also a recessive lethal factor; because of this latter effect of the character no males of this race exist, and the females of the race are never pure but hybrid. Every female with notch wings bred to a wild male, will produce in equal numbers notch winged daughters and daughters with normal wings. There will be half as many sons as daughters. The explanation of